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intertidal zone; where is it
narrow fringe area between high and low water
whats the most important environmental factor in the intertidal zone
tides
diurnal tides
location has a single high and single low tide
semidiurnal tides
two highs and two lows per day
mixed tides
mixture between diurnal and semidiurnal
_____ of the tide varies from day to day
height
why are the tides in intertidal zones important
duration of exposure and time of exposure
tidal rhythms examples
running of the grunion and spawning of the horseshoe crab
environmental conditions of intertidal zones (3)
temperature, wave action, and salinity
temperature in intertidal zones
the organisms are more subjected to more dramatic variations in temp
wave action in intertidal zones
exerts more influence on organisms and communities; mechanical effect (smashes and tears away); extends scope
salinity in intertidal zones
excessive rain/flooding during low tides; tide pool
adaptations of intertidal organisms
avoiding or minimizing the stresses of daily exposure to air
adaptations to withstand water loss in intertidal zones (3)
mobility, dense aggregations, and close up shop
adaptation of mobility in intertidal zones
mobile animals move into moist cracks, crevices, or burrows
adaptation of dense aggregation in intertidal zones
dense groups of organisms; reduce SA; create cracks and crevicesada
adaptation of closing up shop in intertidal zones
close their shells at low tide
rocky intertidal shores
most densely inhabited; greatest diversity of animals and seaweeds
zonation
horizontal banding of organisms
variations of zonation in intertidal zones
slope, tidal range, and exposure to wave action
broad zones of intertidal zones
gradual slope, greater tidal range, and exposed areas
zone classification (3 zones)
supralittoral fringe, midlittoral zone, and infralittoral fringe
supralittoral fringe
uppermost zone; lower limit is the upper limit of barnacles and extends to the upper limit of snails
midlittoral zone
broadest in scope; upper limit is the upper limit of barnacles and its lower limit is the upper limit of kelps
infralittoral fringe
lowest zone; extends from the lowest low tide to the upper limit of large kelps
2 physical factors for causes of zonation
desiccation and temperature
5 biological factors for causes of zonation
competition, predation, grazing, positive interactions, larval settlement
biological factors of zonation in intertidal zones are ____ complex, ____ subtle, and ____ linked to other factors compared to physical factors
more, more, closely
competition in zonation in intertidal zones
doesn’t occur if there are plenty of resources
what one resource is almost always limited in the intertidal zone
space
Whats the name of the dominant space competitor on open-coast shores
Mytilus californianus
Grazers
Herbivores regulate the upper and lower species in the intertidal; they determine which species
species diversity is much ____ in the atlantic than in the pacific
less
algal grazing has led to the evolution of algal defenses (3)
calcification of tissues, chemical defenses, multiple life stages
positive interactions in intertidal zones (2 examples)
mussel beds provide refuge from physical stresses for many organisms; intertidal seaweeds provide shelter and shade for organisms and extend their range in the upper intertidal
most intertidal organisms have a ______ larval stage
planktonic
large variation in recruitment is drive by what facots and conditions (2)
oceanographic conditions and biological factors
oceanographic conditions (2)
strong wilds that lead to upwelling and surface currents moving offshore, strong spring rains can mean decrease in ‘flushing time’ of the bay (washing larvae out of the area)
biological factors of recruitment in intertidal zones (1)
juvenile rockfish in kelp forests are significant predators of barnacle larvae
tropical rocky intertidal characteristics (6)
high temps (basically constant), high predation, mobile inverts abundant, less competition for space, larger # and diversity of consumers, lots of periwinkles
temperate intertidal characteristics (3)
lots of mussels and barnacles, large body size/residence in high intertidal are important refuges from predators, macroalgae have large impact on community structure
coral reef facts (5)
inshore waters dominated by coral reefs, greatest diversity, ½ calcium in oceans taken up by reefs as CaCo3 (so they remove hella carbon per year from atmosphere), primarily tiny colonial corals
biological factors in intertidal zone
settlement preference, competition for space and food, predation
settlement preferences (2)
soft versus hard, zonation
two types of corals
hermatypic and aheramtypic (build reefs vs dont)
reef building corals in phylum cnidaria (one class, one order)
class anthozoa, order scleractinia (stony corals)
example of zonation settlement preference
barnacle settles middle-upper OR throughout intertidal (cant survive exposure any higher)
nutrition for coral
colonial (polyp), carnivorous animals (use nematocysts), colonies of thousands of individuals (total area for feeding is large), feed only during the night time
competition for space and food in intertidal zone
can exclude organisms in intertidal zone
example of competition for space and food in intertidal zone
barnacles grow larger and faster than others and physically excludes it for space
why are the pacific coasts not entirely M. californianus
predation, habitat, human activity, climate, etc.
meroplankton explains only a tiny amount of corals total food requirements. where does the remainder of their food come from?
symbiotic relationship with zooxanthellae (dinoflagellate)
predation in intertidal zone (what animal)
predatory sea stars
how does a zooxanthellae get food from dinoflagellate
photosynthesis (which is then transferred to the corals)
what do dinoflagellates get in their symbiotic relationship with zooxanthellae
protection (coral with energy) and nutrients
common characteristics of rocky shores (5)
no strong detrital component, lots of suspension feeders, lots of macroorganisms, complex predator-prey and competitor interactions, temperate (similar tidal zonation)
how are zooxanthellae acquired
directly (asexual budding) or indirectly from the environment
tide pools characteristics (3)
salinity changes, temperature changes, and oxygen depletion
how do corals get color
zooxanthellae (they feed by photosynthesis and have color)
how does coral get bleached
zooxanthellae get stressed and dip
growth and calcification of a coral colony (requirements)
light, growth rate varies by species, growth rate varies by location
intertidal fishes (4)
clingfish, gobies, blennies, sculpins (and other visitors)
how does light help with growth and calcification of coral reefs
zooxanthellae increase the calcification of corals and growth rate of colonies
how does growth rate vary by SPECIES in coral reefs
branched corals grow more rapidly than massive or mound shaped corals
how does growth rate vary by LOCATION in coral reefs
corals at a greater depth are more thin and fragile
why are corals at a greater depth more thin and fragile
greater depth = cold and dark (not ideal)
reproduction in coral reefs
asexually and sexually
sexual reproduction in coral reefs
results in free swimming PLANULA larvae
asexual reproduction in coral reefs
new individuals bud off the parent
characteristics of planula larvae (3)
sexually mature at 7-10 years, hermaphroditic, seasonal broadcast spawners
recruitment in coral reefs
planulae disperse in open water few days/weeks until settle to start new colony, and the rate of recruitment is variable in time and space
why is the rate of recruitment in coral reefs variable in time and space
mortality of new colonies is thought to be high
SIX major physical factors limit coral reef development
temperature, sunlight, depth, salinity, sedimentation, exposure to air
reef distribution (temperature)
no reefs below 18 C, optimal at 23-25 C (upwelling makes areas cold)
reef distribution (depth)
do not develop it waters deeper than 50-70m, optimal at 25m
reef distribution (light)
sufficient light for photosynthesis by symbiotic zooxanthellae
reef distribution (salinity)
corals are marine organisms and intolerant of salinity change
reef distribution (sedimentation)
can clog feeding structures and smother the corals, turbidity reduces light
3 types of reefs
Fringing, barrier, and atolls
Barrier reef
Distance between land and reef
Atoll reef
Circular/horseshoe shaped reef far from land usually deep
Subsidence theory
Fringing reef → barrier → atoll (island sinking)
Why does an island sink (subsidence theory)
It used to be a volcano, cooled down, and got denser
how can an island sinking affect the coral around it
If the island sinks faster than the reef builds on top of itself, it will de
Enewetok
In the Marshall Islands, an atoll reef
Marshall Islands
100 B.C. And important in ww1 and ww2; ships were refueled there
Donation on coral reefs
Great diversity of life
Fringing reef
Reef surrounds land
zonation on coral reefs (diagram)
Outer seaward slope 2 windward reef margin 3. Algal ridge 4.reef flat
Seaward beach of island 6. Lagoon beach of island 7. Lagoon floor
Lagoon reefs 9. Leeward reef flat 10. Leeward reef margin 11. leeward reef slope

Depth zonation in coral
Reefs on surface, branching on top of slope, massive corals at bottom of slope
Competition in coral reefs
Competition for light and space (lack of space in reefs) to grow higher and wider by growing faster
Exploitative competition n coral reefs
Up right, rapid growing corals grow over encrusting massive corals; slow growing corals have adapted to tolerate shade
Interference competition
Slow growing corals extend digestive filaments and kill adjacent competing species