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Soil
layer of chemically and biologically altered material over parental material or bedrock
Parental material
Rock or material that soil is created from*
Leaching
Precipitation/groundwater removes some substances from soil by dissolving them and moving them down to lower layers
Weathering
Physical and chemical alteration of rock material near Earth’s surface; occurs when surface water penetrates parent material
Water potential
measure of the potential energy of the water, indicating its tendency to move from one area to another
Dependent on: gravity, pressure, osmotic potential, matrix potential
Matrix potential
Energy by the electric attractive forces between the water molecules and the matrix of soil particles
Field capacity
Maximum amount of water held by soil particles against the force of gravity (gravity = attractive forces in soil)
Wilting point
Water potential at which most plants can no longer retrieve water from the soil; point at which water potential in plant is unable to counteract matrix potential and absorb water, despite the presence of water
Loam
mixture of sand, silt and clay; the best soil for growing plants (40% sand, 40% silt, 20% clay)
Cation exchange capacity
ability of a soil to retain cations; determines fertility of a soil
Transpiration
Process by which leaves can generate water potential as water evaporates from the surfaces of leaf cells into the air spaces within the leaves
Stomata
perforations that allow for water to leave the plant, but also for exchange of O2 and CO2
Cohesion-tension theory
explanation of the mechanism of water movement from roots to leaves due to water cohesion and water tension
Photosynthesis
two part chemical reaction: light reaction, Calvin cycle
Calvin Cycle
ATP and NADPH are used to fix carbon dioxide and produce chemical energy as sugars
C3 photosynthesis
widespread photosynthetic pathway occurring in mesophyll cells; in algae, aquatic plants, and most terrestrial plants
(CO2 + RuBP = G3P as main sugar)
C4 photosynthesis
photosynthesis steps are separated into space (Co2 assimilation in mesophyll cells, rest in bundle sheath cells); in some herbaceous plants in hot and dry climates (sugarcane, sorghum, corn, etc)
CAM photosynthesis
Same biochemical pathway than C4, but separated in time rather than space (stomata open at night, close during day); happens entirely in the mesophyll cell; some succulents from water stressed environments (cacti, pineapple, etc)
Mesophyll cells
exclusively where C3 photosynthesis occurs
Bundle sheath cells
where part of calvin cycle happens in C4
Malic acid
form that carbon is stored as in C4 photosynthesis; is transported to budle sheath cells and is broken int CO2 and pyruvate
Radiation
gain and loss of heat from sun and organisms
Conduction
gain and loss of heat between objects
Convection
gain and loss of heat by movement of gases and liquids
Evaporation
gain and loss of heat by energy used to evaporate a liquid
Thermal inertia
property of a material to resist changes in temperature by both storing and delaying the transmission of heat (smaller organism=lower thermal inertia)
Blood shunting
blood vessels can be shut off using precapillary sphincters, directing the blood back to the heart rather than extremities to prevent heat loss
Temporal environment variation
-predictable variation: day/night, seasons, etc
-unpredictable variation: quick weather changes/events
Spatial environmental variation
-large scale: climate, topography, soil type
-small scale: structure of plants, animal activity, soil composition
Phenotypic trade off
organisms can adapt only to a specific environment; one phenotype experiences high fitness in one environment, another phenotype experiences higher fitness in another environment
Phenotypic plasticity
response to environmental variation by evolving morphology, physiology, behavior, growth, development, reproduction, etc.
Environmental cues
Cues that trigger phenotypic elasticity
biotic cues
abiotic cues
Biotic cues
“enemies” (predators, herbivores, parasites, pathogens), competitors, mates
Abiotic cues
temperature, water availability, salinity oxygen, photoperiod, etc
Migration
seasonal movement of animals from one region to another
Diapause
a period of suspended development in an insect or other invertebrate, especially during unfavorable environmental conditions*
Endotherms
produce own body heat (mammals, birds)
Ectotherms
body heat is taken from environment
Hibernation
endotherms reduce heart rate and body temperature*
Torpor
brief periods of dormancy in birds and mammals to cope with low temperatures
Aestivation
shutting down of metabolic process in snails, desert tortoises, and crocodiles in hot and dry summers
Optimal foraging theory
model that describes foraging behavior considering costs and benefits of different foraging strategies
Four possible responses to food variation:
central place foraging
risk-sensitive foraging
optimal diet composition
diet mixing
Central place foraging
requires acquired food to be brought to a central place
Risk-sensitive foraging
incorporates the risk of predation when making foraging decisions
Optimal diet composition
choice from a range of food items to weigh trade-off of energy vs handling time, and abundance of prey
Life history
schedule of growth, development, reproduction, and longevity
Life history traits
time to sexual maturity
fecundity
parity
parental investment
longevity
Fecundity
number of offspring per reproductive period
Parity
number of reproduction events
Parental investment
time and energy given to offspring
Longevity
life expectancy
Competitors
plants that are efficient at resource capture and utilization, often exhibiting large root systems or tall growth to maximize resource acquisition in environments with abundant resources but minimal threats
Stress tolerators
species adapted to survive in harsh conditions with limited resources; typically grow slowly, have slow metabolism, and invest in mechanisms for maintenance and survival, such as detoxification or defenses
Ruderals
hort-lived and fast-growing plants that focus on rapid seed production and dispersal to quickly colonize disturbed areas
Metamorphosis
process of transformation from an immature form to an adult form in two or more distinct stages, involving significant physical changes in body structure, physiology, and behavior
Semelparity
One reproductive event per lifetime
Iteroparity
multiple reproductive events per lifetime
Senescence
increased likelihood of dying as aging occurs
Vegetative reproduction
somatic tissues of the parent produce clones
Parthenogenesis
Embryos are produced from germ lines but without fertilization; normally in egg-producing organisms; offspring can be clones (no meiosis) or genetically variable (meiosis)
Simultaneous hermaphrodites
have both male and female reproductive organs; can sometimes self fertilize
Sequential hermaphrodites
sex changes throughout lifespan
Indeterminate growth
ongoing process of growth in an organism, which adds new cells and allows the organism to continue to increase in size, produce new parts, or grow after reaching sexual maturity
Isogamy
Organisms produce similar gametes in both males/females
Anisogamy
Each parent produces a different gamete (i.e. sperm/eggs)
Hermaphroditism
Organism produces both large and small gametes
Sexual dimorphism
observable differences in physical and behavioral traits between males and females of the same species
Haplodiploidy
sex-determination system where males are haploid (developing from unfertilized eggs) and females are diploid (developing from fertilized eggs)
Hymenopterans
bees, wasps, ants, etc.
Frequency dependent selection
if all sexxes have similar fertility, the rarer sex in the population is favored over the commonest sex by natural selection
Primary sexual traits
traits related to fertilization (eg. gonads)
Secondary sexual traits
traits not related directly to fertilization (eg. body size, ornaments, armaments, behaviors)
Mating systems
methods of mating interactions
promiscuity
polygamy
monogamy
extra pair copulation
Promiscuity
individuals mate with multiple partners and do not create a lasting social bond
Polygamy
individuals form long-term social bonds with multiple mates
polyandry
polygyny
Polyandry
females mating with multiple males; males offer a reward for mating (prey or spermatiphore)
Polygyny
males mating with multiple females
harem
lek
territory
Monogamy
species form a social bond at least while rearing offspring
Extra pair copulation
mating outside of a recognized pair bond, often in socially monogamous species
Harem
a polygynous mating system where one male defends a group of females and their offspring for the purpose of exclusive or favored mating
Lek
an area where males of a particular species gather to display and attract females for mating, without any other resources being offered
Territory
a specific area that an animal aggressively defends from conspecifics (individuals of the same species) to secure access to resources and opportunities for mating
Sex roles
role of each sex in reproductive interactions
Material benefit
food, quality of territory, location to lay eggs, etc
Non-material benefit
good genes hypothesis, good health hypothesis
Runaway sexual selection
process where mate choice and trait elaboration create a positive feedback loop, leading to the evolution of increasingly exaggerated ornaments or behaviors that may be costly to survival
Handicap
individuals with high fitness can afford to produce costly, elaborate, or disadvantageous signals to honestly communicate their genetic quality to potential mates or rivals
Honest trait
biological signal that reliably conveys accurate information about an individual's quality or condition to other organisms (ie. vibrant feathers in male peacock)
Sexual conflict
mating partners behave according to their own self-interest leading to conflict
Mate guarding
one individual prevents the other from participating in extra-pair copulation
Copulatory plugs
plugs make future mating impossible
Infanticide
new dominant male kills offspring of last male to increase own fitness
Traumatic insemination
physical harm to females upon mating to ensure the female lives shorter and lays fewer eggs the more males they mate with
Good genes hypothesis
the idea that females choose mates exhibiting certain traits because these traits signal underlying superior genetic quality, which increases the offspring's chances of survival and reproductive success
Good health hypothesis
individuals choose mates based on traits that honestly signal superior genetic quality
What materials does soil contain?
minerals, decomposing organic matter, community of microorganisms, invertebrates and insects
What does soil composition depend on?
parental material, climate, vegetation, age, local topography
What impacts water potential?
gravity, pressure, osmotic potential, matrix potential
What is the soil composition of dry climates? Why?
Little rainfall causes slow parent material breakdown and sparse plant growth; little organic material is added to soil; shallow soils with close-lying bedrock layer