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Cutaneous respiration
diffusion through the membrane - a process where gas exchange occurs through the skin. Pros: minimal energy. Cons: you have to be small and simple
Tracheal system (arthropod system)
Series of branching tubes
Can be ventilated with movement
Direct transport to cell
Independent of a circulatory system
First step!
Countercurrent flow
a mechanism where two fluids (liquids or gases) move in opposite directions across a boundary to maximize the transfer of heat or a substance, like oxygen
CONCEPT: Selection differs in aquatic versus terrestrial systems
Selection differs between aquatic and terrestrial systems due to the contrasting physical and chemical environments, which shape traits like feeding, attachment, and life history strategies
CONCEPT: Smaller organisms can acquire oxygen via simpler, more direct mechanisms
Smaller organisms can effectively respire via passive diffusion or cutaneous
CONCEPT: Countercurrent flow maximizes gas exchange
Countercurrent flow maximizes gas exchange by having fluids move in opposite directions, which maintains a constant concentration gradient that allows for more efficient transfer of a substance like oxygen from water to blood in fish gills.
CONCEPT: Mammals have a highly inefficient respiratory system
Air exits where enters; bidirectional
Dead air in passageways where gas exchange does not occur
Passive gas exchange between capillaries and alveoli, active in the diaphragm
CONCEPT: Birds have an efficient, unidirectional system adapted for flight
Air enters through the trachea directly into air sac - then to the lungs, then again to the second air sac, then exits the body
2 cycles of inhalation/exhalation
CONCEPT: Efficiency in the respiratory system is related to metabolic rate
The body needs to supply cells with the oxygen required for metabolic processes like cellular respiration. A higher metabolic rate requires more oxygen, which in turn necessitates a more efficient respiratory system to meet the demand
Gastrovascular cavity
Central digestive/circulatory compartment in simple animals
One opening
Diffusion
Small, aquatic animals
Open circulatory system
Lacks blood vessels, capillaries
Has hemolymph (fluid) and hemocoel (blood sinus)
Water, proteins, salts, phagocytic cells
20-40% volume!
Simplest in terrestrial organisms
BATHED IN BLOOD
Single closed circulatory system
Distinct blood vessels, capillaries
Hydrostatic pressure - what differentiates closed vs open circulatory systems - blood can move fast! More efficient
Failsafe valves
5-10% volume!
Heart = pumps one time in circuit
Sinus venosus - holding chamber before atrium
Blood pressure highest before ventricle
Conus or Bulbous arteriosus = holding chamber after ventricle
Slow and low metabolism and blood pressure
Double closed circulatory system
Distinct blood vessels, capillaries
Hydrostatic pressure - what differentiates closed vs open circulatory systems - blood can move fast! More efficient
Failsafe valves
5-10% volume!
Differential pumping pressure
Larger body size
Pulmonary circuit - low pressure
Systemic circuit - high pressure
Importance: endothermy, terrestrial ecology, body size
Diastole
relaxation (chambers fill)
Systole
contraction (blood pumped)
Main anatomy of mammal heart
Left atrium (oxygenated)
Left ventricle (oxygenated)
Tricuspid valve
Right atrium (deoxygenated)
Right ventricle (deoxygenated)
Bicuspid valve
Aorta
Hemoglobin
Oxygen transportation
All kingdoms (common), not all organisms
Vertebrates
Binds 4 oxygen
Confined to RBCs
Terrestrial environments
Red
Hemocyanin
Oxygen transportation
2nd most common
Cephalopods (Mollusca) & Crustaceans (Arthropoda)
Hemolymph
2 copper (Cu) binds to 1 oxygen
Free-floating protein, high densities
Cold, low oxygen environments
Efficacy decreases with acidification
Colorless to blue
CONCEPT: Circulation is important for transporting gas, nutrients, and waste
Circulation delivers oxygen from the lungs and nutrients from the digestive system to all cells, while also removing waste products like carbon dioxide, which is then taken to the lungs to be exhaled.
CONCEPT: Cellular respiration in larger animals requires respiratory pigments, with oxygen saturation variable throughout the circulatory system
Larger animals require respiratory pigments like hemoglobin to transport enough oxygen for cellular respiration
CONCEPT: Differences in the efficiency of the pump systems in closed v. open systems are tightly correlated with metabolism & ecology
Open system - small, simple organisms
Closed system - single pump = slower metabolism, double pump = larger bodies and higher metabolism
CONCEPT: Respiratory pigments (hemoglobin & hemocyanin) facilitate gas exchange & also are correlated with ecology
Respiratory pigments like hemoglobin and hemocyanin facilitate gas exchange by dramatically increasing the blood's oxygen-carrying capacity beyond what can be dissolved in plasma alone. Their structure and function are strongly correlated with an animal's ecology, adapting to environmental factors like oxygen availability, metabolic needs, and habitat type.
CONCEPT: Metabolic rate is associated with body size and endothermy
For endotherms (warm-blooded animals), smaller animals have a higher metabolic rate per unit of mass compared to larger animals. This is because their larger surface-area-to-volume ratio leads to faster heat loss, requiring a higher metabolic rate to maintain a stable internal temperature. Additionally, endothermy as a whole results in a much higher metabolic rate than that of an ectotherm (cold-blooded) of the same size, because endotherms must expend energy to generate their own body heat.
CONCEPT: More specialized circulatory systems are important for the evolution of endothermy, terrestrial ecology, and larger body sizes
Specialized circulatory systems were crucial for the evolution of endothermy, terrestrial ecology, and larger body sizes because they enabled the efficient transport of oxygen and nutrients necessary to meet higher metabolic demands and overcome the limitations of simple diffusion over greater distances - and 4 chambers with specialization means no deoxygenated blood left circulating around body.
External respiration
Occurs as a function of partial pressure differences in oxygen and carbon dioxide between the alveoli and the blood in the pulmonary capillary
Internal respiration
Occurs as a function of partial pressure differences in oxygen and carbon dioxide between tissue and the blood in the capillary, which are opposite of those present at the respiratory membrane