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Ethology
the study of how evolutionary processes shape inherited behaviours and the ways animals respond to specific stimuli
Behaviour
an animal’s response to a stimulus
Proximate cause
What was the stimulus to cause the behaviour?
Ultimate cause
How does the behaviour help the animal survive and reproduce?
Fixed Action Patterns
a sequence of unlearned acts directly linked to a stimulus
Migration
the regular, long-distance change in location
Signal
a stimulus generated and transmitted from one animal to another
Pheromones
chemicals emitted by members of a species that can affect other members of the same species
Stimulus response chains
when a response to a stimulus serves as the next stimulus for a behaviour
Directed movements
movements towards or away from a stimulus
Kinesis
a change in the rate of movement or the frequency of turning movements in response to a stimulus, non-directional
Taxis
directional movement towards (positive) or away from (negative) a stimulus
Phototaxis
Movement in response to light
Chemotaxis
movement in response to chemical signals
Geotaxis
movement in response to gravity
Learning
a modification of behaviour based on specific experiences
Imprinting
a long lasting behavioural response to an individual
Spatial learning
establishing memories based upon the spatial structure of the animal’s surroundings
Associative learning
the ability to associate one environmental feature with another
Social learning
learning through observations and imitations of the observed behaviours
Foraging
food obtaining behaviour that involves searching for and exploiting food resources
Mating behaviours
animals can be monogamous or polygamous
Atruism
selfless behaviour that benefits others, often at a cost to oneself
Phototropism
a directional response that allows plants to grow towards a source of light
Photoperiodism
allows plants to develop in response to day length: plants flower at only certain times of the year
Physical defences
thorns, trichomes, etc
Chemical defences
production of toxic or distasteful compounds
Ecosystem
the sum of all the organisms living in a given area and the abiotic factors they interact with
Biotic
living, or once living, components of an environment
Abiotic
nonliving (physical and chemical properties) of the environment
1st Law of Thermodynamics
energy can neither be created nor destroyed, only transferred
Law of Conservation of Mass
chemical elements are continually recycled in the environment
2nd Law of Thermodynamics
exchanges of energy increase the entropy of the universe
Metabolic rate
the total amount of energy an animal uses in a unit of time
Endotherms
use thermal energy from metabolism to maintain body temperatures
Ectotherms
use external sources to regulate their body temperatures
Heterotrophs
rely on autotrophs because they cannot make their own food
Autotrophs
use light energy to synthesise organic compounds
Primary consumers
herbivores
Secondary consumers
carnivores that eat herbivores
Tertiary consumers
carnivores that eat carnivores
Decomposers
get energy from detritus (nonliving organic materials)
Food chain
the transfer of food energy up trophic levels
Food webs
Linked food chains
Primary production
the amount of light energy that is converted to chemical energy
Gross primary production (GPP)
total primary production in an ecosystem
Net primary production (NPP)
the GPP minus the energy used by the primary producers for respiration (Ra)
Secondary production
the amount of chemical energy in a consumer’s food that is converted to new biomass
10% rule
the transfer of energy between trophic levels is around 10% efficiency
Water cycle
water is essential for all life and influences the rate of ecosystem processes
Carbon cycle
carbon is essential for life and required in the formation of organic compounds
Nitrogen cycle
nitrogen is important for the formation of amino acids, proteins, and nucleic acids
Phosphorous cycle
phosphorous is important for the formation of nucleic acids, phospholipids and ATP
Population
a group of individuals of the same species living in an area
Population ecology
analyses the factors that affect population size and how and why it changes over time
Dispersion
the pattern of spacing among individuals within a population
Clumped
individuals gather in patches
Uniform
evenly spaced individuals in a population (can be due to territoriality)
Random
unpredictable spacing, not common
Demography
the study of the vital statistics of populations and how they change over time
Life table
an age-specific summary of the survival pattern of a population, represented by a survivorship curve
Type I curve
low death rate during early/middle life and high death rate later in life
Type II curve
constant death rate over the lifespan of the organism
Type III curve
high death rate early in life and lower death rate for those that survive early life
Per capita rate of increase formula
dN/dt = B - D, where dN/dt is the change in population size, B is the birth rate, and D is the death rate per capita.
Exponential growth model
a population is living under ideal conditions, population grows rapidly
Exponential growth formula
dN/dt = rmaxN, where rmax is the maximum per capita growth rate, and N is the current population size (J shaped curve
Logistic growth model
the per capita rate of increase approaches zero as the population size nears its carrying capacity
Logistic growth formula
dN/dt = rmaxN(K-N/K) where K is the carrying capacity, and N is the current population size and rmax is the per capita growth rate
Life history
the traits that affect an organism’s schedule of reproduction and survival, including age at first reproduction, frequency of reproduction, and number of offspring produced
K-selection (density dependent)
selection for life history traits that are sensitive to population density, seen in high density populations that are close to carrying capacity
R-selected (density independent)
selection for life history traits that maximise reproductive success, seen in low density populations with little competition
Density-dependent regulation
as a population increases, factors can slow or stop growth by decreasing birth rate and increasing death rate (competition, predation, toxic wastes, territoriality, disease, intrinsic factors)
Density-independent regulation
factors that exert their influence on population size, but the birth/death rate remains the same (weather, climate, natural disasters)
Community
a group of populations of different species living closely and capable of interacting
Habitat
a place or part of an ecosystem occupied by an organism
Ecological niche
the role and position a species has in its environment
Fundamental niche
the niche potentially occupied by the species if there were no limiting factors (predators, competitors, etc)
Realized niche
the portion of the fundamental niche the species actually occupies
Interspecific interactions
interactions of individuals from one species with individuals of another species
Competition
-/- relationship where individuals of different species compete for limited resources
Competitive exclusion principle
two species competing for the same resource cannot coexist permanently
Niche partitioning
natural selection drives competing species into different patterns of resource use, or different niches
Predation
+/- relationship where one species (predator) kills and eats the other species (prey)
Cryptic coloration
camouflage
Batesian mimicry
harmless species mimics a harmful one
Mullerian mimicry
two or more bad-tasting species resemble each other
Herbivory
+/- relationship where one organism eats part of a plant or alga
Symbiosis
when 2 or more species live in direct contact with one another
Parasitism
(+/-) when one organism (parasite) derives nourishment from another (host)
Mutualism
(+/+) when both organisms benefit from the relationship
Commensalism
(+/0) when one organism benefits and the other is neither harmed nor benefited
Facilitation
(+/+ or 0/+) when one species has a positive effect on the survival and reproduction of another without intimate association of symbiosis
Species diversity (biodiversity)
the variety of different organisms within a community
Species richness
the number of different species
Relative abundance
the proportion each species represents of all the individuals in the community
Simpson’s diversity index
calculates diversity based on species richness and relative abundance, high diversity index means high biodiversity; low diversity index means low biodiversity
Simpson’s diversity index formula
Diversity index = 1-∑(n/N)2 where n is the number of individuals of a species and N is the total number of individuals of all species
Invasive species
non-native species that disrupt ecosystems and outcompete native species
Keystone species
not usually abundant, but other species in an ecosystem rely on them because of their important ecological niches