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Species
organisms that can produce fertile offspring
Population
all of the individuals of a single species that interact/breed
The unit at which evolution acts
Community
all of the populations of living things that interact in a place
Ecosystem
biological community of interacting organisms and their physical environment
Abiotic and biotic factors
Abiotic Factors that Define Ecosystems
Climate
-Averages and seasonality
Geology
-Bedrock, water, soils, topography
Disturbance
-Natural, human
-Scale and frequency
Biotic Factors that Define Ecosystems
Biodiversity
Species Interactions (Predator-Prey, Mutualism, Commensalism, etc.)
Presence of Invasive or Keystone Species
Ecosystems cannot be defined by
size
boundaries
Mt. St. Helens
Volcano that is part of Cascade Mountains that erupted in Washington state causing massive damage in 1980
-in temperate rainforest biome
Pre-Eruption Mt. St. Helens Ecosystem
Old Growth forest
-Never been logged
-Diverse forests and species
Rich soils
-Stimulated plant growth with lots of available nutrients
Heterogenous ecosystem that changed with elevation
temperate rainforest
Post-Eruption Mt. St. Helens Ecosystem
Forests scorched up to 17 miles from mt. st helens
Pumice Plain covered in ash so low nutrient levels in soil
Why do Mountain ecosystems have such high biodiversity?
As elevation increases, temp, O levels, precipitation levels, and biodiversity change and the ecosystems change
Elevation and Latitude: Differences in elevation in an area allow multiple biome
Effect of the eruption of MSH being lateral?
Northern slopes decimated, but southern slopes relatively unaffected
Scorch Zone
outermost forest
-defoliated trees
-surviving ground vegetation due to snow on the ground
debris avalanche
riverbeds where melt water and mud was washed in
Pumice Plane/Pyroclastic Flow
closest to blast zone
-Feet of volcanic ash (tephra/rock dust)
-No life
-small portion of the affected landscape
Blowdown Zone
largest zone
-Blast knocked over trees
-Not much surviving ground vegetation
Spirit Lake
flooded with muddy ash water and logs
-Murky water, limited visibility
What affects the order of reestablishment?
What can reach the area first?
-By wind(spores/seeds)
-What can survive there?
The first to survive there will be the first to reestablish
Plant Dispersal
how seeds/spores are spread from the parent plant
-Purpose: increases chance of survival by limiting competition for resources (soil nutrients..)
-Methods: wind, animal, water, gravity
Wind Dispersal
A method of dispersal where lightweight seeds catch the wind
-Long distances, random
Animal Dispersal
Seeds have hooks attached to them that attach to fur. Other seeds are eaten and pass through the digestive tract. can be carried far distances but varies
Water Dispersal
Seeds fall into the water and float away
-Splashes or long distance
Gravity Dispersal
Heavy nuts fall to the ground and roll
Example: Ballistic Dispersal
Ballistic Dispersal
plant catapults/ejects seeds
Ex. lupine
Seed Coat
protection and can impact dispersal
-Thickness varies (example. Walnuts vs. sunflower seeds)
-Thicker seed coats offer better protection from animals and dehydration
-Thinner seed coats don't cost as much energy to produce, can migrate fast

Cotyledon
food storage of fats, proteins, and oils
-Can be big or small
-Bigger cotyledons can offer more storage and last longer before germination (sprouting), higher survival rates, can grow quickly to overtop competitors
-Smaller cotyledons allow the seed to disperse farther and faster(lighter), takes less energy to produce (plant can produce more seeds)

Evolutionary Tradeoff
a trait increases fitness in one condition, while decreasing fitness in another
-explains why large-seeded plants produce relatively few seeds
per capita survivorship
probability of survival for any one seed
-decreases with the size of the seed (big seeds take more energy to produce but are more likely to survive)
Fitness
the ability to pass on genes to subsequent generations
Macronutrients
needed in large quantities to make nucleic acids and amino acids
-Primary (N, P, and K) and secondary nutrients
-Examples: Nitrogen, Phosphorous
Micronutrients
needed in small quantities, coenzymes
-Come from soil
-Example: Zinc
Where do plants get C?
air

Where do plants get H?
soil
Where do plants get O?
soil and air

What are plants made of?
50-75% H2O
-C, H, O (cellulose is C6H10O5, glucose is C6H12O6)
-N (big component of DNA and proteins)
-P (big component of phosphate backbone of DNA)
-Also some Magnesium (makes chlorophyll)
Nitrogen Cycle
(1) nitrogen fixation
-Nitrogen gas to ammonia
-carried out by bacteria typically
(2) nitrification
-ammonia to nirates and nitrites
-carried out by nitrifying bacteria typically
(3) denitrification
-nirates and nitrites to nitrogen gas and nitrous oxide

Tephra
-rock dust
-MSH soil
-missing nitrates
Importance of Nitrogen
-used to make proteins responsible for muscles,cellular communications and enzymes
-used for DNA's nitrogenous bases
-used to make chlorophyll
Protein Folding Steps
Primary structure, Secondary Structure, Tertiary Structure, Quaternary Structure
Proteins
Complex molecules made of amino acids
-Synthesized using ribosomes and an mRNA template (translation)
-Ex. digestive enzymes, helicase, muscle fibers
Primary structure: straight chain of amino acids, no folding, held together by peptide bonds
Secondary Structure:
Crimped (Beta Sheet)
Curled (alpha helix)
Tertiary Structure: multiple folds and bends of a single amino acid chain
Quaternary Structure: multiple tertiary structures
Primary structure
straight chain of amino acids, no folding, held together by peptide bonds
Peptide Bond
C-N polar covalent bond (e- shared) connecting AA (C from carboxyl group and N from amino group
-primary structure
Secondary Structure
Crimped (Beta Sheet)
Curled (alpha helix)
-H-bonds between Carboxyl and amino groups of AAs
Tertiary Structure
-multiple folds and bends of a single amino acid chain
-R- groups determine 3D structure
-IMFs, ionic bonds, and sometimes disulfide bridges determine exact folding pattern
quarternary structure
multiple tertiary structures
What can determine proteins shape?
-bonds, IMFs
-Can denature when too hot or wrong pH and lose its function
-mutations in an amino acid and that will change its function
Ionic Bond
-donate/take e-
-big differences in electronegativity in 2 molecules
-Permanent attraction between whole charged molecules (+1/-1)
-Anions: negative charge, extra electron, gained an e-
-Cations: positive charge, missing e-, lost an e-
Ex. NaCl
Covalent Bond
Share e-
-Low energy state
Ex. HCl
Polar Covalent Bond
E -shared unequally
-partial charge associated with a bond

Nonpolar Covalent Bond
E- shared equally

IMFs (intermolecular forces)
electrostatic charge attractions between molecules that can vary in permanence and strength
-H-bonds, Dipole-Dipole
Hydrogen Bond
strong temporary attraction between partial positive charges of H and partial negative N,O, F, Ex. H2O
-connects nitrogenous bases in DNA
Electronegativity
how tightly an atom holds onto its e-

components of amino acids
Amino Group
Carboxyl Group
R Group (determines AA interactions and protein shape)
What is the Limiting Resource in MSH soil?
Nitrogen
Liebeg's Law of the Minimum
Assumptions:
1) 1 limiting resource at a time
2) Too much of a nutrient isn't harmful
3) Ignore size limitations for organisms
-Can apply to individuals and populations: (states that they will grow only up to the point it runs out of a vital resource, even if there are surpluses of other resources)
-Only works with organisms with indeterminate growth
Indeterminate Growth
-don't have a maximum size, they grow until they die
Ex. trees, goldfish, lobsters, some snakes, etc.
Lupine
-1st to reestablish in MSH (pioneer species)
-improved the pumice plain by decomposing and enriching the soil, facilitating colonization of other plants
- has a mutualistic relationship with N-fixing bacteria that grows in its roots (plants get usable N and bacteria get habitat, glucose and other photosynthesis products)
Haber-Bosch Process
artificial way of producing N
-Caused huge agricultural revolution in the early 1900s
Types of Species Interactions
Mutualism(+/+), Predator/Prey(+/-), Parasitism(+/-), Herbivory(+/-), Commensalism(+/0), Altruism(-/+), Facilitation(-/+), Competition(-/-)

Mutualism (+/+)
Both species benefit from the interaction
Symbiotic Relationship
Live their lives in close physical proximity and evolved together (does not necessarily mean either organism is helped, harmed, or neither)
Ex. Clownfish and anemone (clownfish gets shelter and anemone is unaffected)
Ex. Mistletoe is a plant who sends root-like structures into the bark of a host tree like an oak in order to intercept and absorb the sugars and nutrients from its host. [Mistletoe and Oak trees]
Ex. Tick and dog (tick gets blood, dog is harmed)
Altruism
(-/+)one organism acts to increase the fitness of another organism at a cost to itself (decreasing its fitness)
Ex. Dog raises a litter of kittens
Facilitation
(-/+) One organism changes the environment, leading to its own eventual replacement
-Typically occurs in plants
-drives succession
Ex. Lupine improved the pumice plain by decomposing and enriching the soil allowing other species to colonize and take over
Ex. Blackberries and Hawthorns
Kin Selection
an organism increases the fecundity of a relative at a cost to its own reproductive capacity
-mutualistic
-Ants and bees
Competition
(-/-) limiting the other's potential
Types: Interspecific, Intraspecific
Interspecific Competition
competition between individuals of different species

Intraspecific Competition
competition between individuals of the same species

How to Plants Facilitate Each Others Growth?
Increased soil moisture
Soil building
Temperature and humidity regulation
Attracting pollinators
Soil Chemistry Alteration (N-Fixers like Lupine)
Succession
process of development that over time, gradually and predictably changes the biological community
Stages: Pioneer Stages, Intermediate Stages, Climax Community
Types: primary and secondary
-driven by facilitation
Pioneer Stages of Succession
1. Bare Rock
2. Lichen (can break down rock for nutrients, decompose and create soil)
3. Small annual plants, moss
4. Perennial herbs, grasses (stabilize soil)
Intermediate Stages of Succession
Shrubs, shade intolerant trees
Climax Community
stage that will persist as astatic ecosystem and will continuously regenerate itself until a disturbance
-Characterized by shade-tolerant trees (can grow without a lot of light, can grow in shade under established trees)

primary succession
succession starting with bare rock
Ex. succession beginning on mountain tops/granite outcrops, glacier retreats, parking lots,

secondary succession
Succession starting with soil
Ex. MSH

Mt. St. Helens Succession
secondary succession
Pioneer Stages
1. Lupine
2. Fireweed/Pearly Everlasting
Intermediate
3. Willows, Red Alder
Climax
4. Noble Fir, Douglas Fir, Red Cedar
What changes with Succession?
1. Complexity
2.Biodiversity
3.Soil Complexity
Complexity
habitat diversity
Types: vertical and horizontal
Increasing complexity —> increases habitat diversity —-> increases biodiversity
Vertical Complexity
Layers of a Forest
1.Soil
2.Forest floor
3.Understory/ Shrub Layer
4.Sub-canopy
5.Canopy
Horizontal Complexity
patchiness (more patches, more diversity, more stages of succession —> more horizontal complexity)
-increased by disturbances
Species Richness
the number of different species
Simpsons' Biodiversity of Index (S')
identifies relative abundance of species
0
Disturbance Regime
refers to the size(scale), type, and frequency of disturbances
Ex. MSH Disturbance Regime: volcanic disturbances (infrequent and large scale eruptions)
Dynamic Equilibrium
an ecosystem in a constant state of flux due to disturbance and succession over times
-Contains patches of different successional stages
-High biodiversity
-High horizontal complexity, varying vertical complexity
Ex. South slope of MSH
Ecosystem Services
economic benefits to humans provided by the natural environment and from healthy ecosystems
Ex. erosion control, pharmaceuticals, coastal hurricane protection

Why are some places more biodiverse than others?
1. Time (succession and Dynamic Equilibrium)
2. Ecosystem Complexity (Horizontal, vertical, topographic)
3. More energy (equator v. poles)
4. Environmental Harshness (Limiting Factors)
A plant that lives in a very dry place where suitable conditions for germination happen infrequently would most likely have evolved to have a _________ seed coat and a ___________ cotyledon.
thick , large
The plants that will be able to reach the interior of the pumice plain first are likely to be seeds with __________ cotyledons and have __________ dispersed seeds.
small , wind
Give an example of a species that will likely be the last to reach the interior of the pumice plain.
Cedar Trees
Charge of P in Phosphoric Acid (H3PO4) where P is central atom connected to 3 OH- groups and one O atom?
partial positive

If all of the hydrogen bonds in a protein were broken, which levels of protein structure would be altered?
Secondary structure
Tertiary structure
Quaternary structure
If 2 atoms share electrons in a covalent bond and their electronegativity difference is 0.3, what does that mean?
They do not share electrons evenly, but it's close enough that we consider the bond to be non-polar
Autotroph
get carbon from inorganic sources like CO2
Heterotroph
get carbon from organic sources (other organisms)like glucose
Are carnivorous plants autotrophs or heterotrophs?
autotrophs
What increases with succession?
Soil Carbon and Nitrogen
Biodiversity
Vertical Complexity
Resistance
Resilience
Shade-tolerant trees
Horizontal Complexity
Compare complexity of permaculture farm to organic farm.
Permaculture farm has high horizontal complexity and high vertical complexity compared to the organic farm
Biomass
the total weight of all the living things in an area
Ecosystem Resistance
ability to prevent impacts from disturbance (less impact of disturbances)
Increase in species richness/biodiversity/complexity —-> higher ecosystem resistance