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Ordovician
§485.4-443.8mya
§Gondwana supercontinent
§Appalachian Mountains form
§Very hot, with cooling towards the end
§High CO2
§Colonization of plants on land
§Ordovician-Silurian extinction
§Potentially caused by plants
Plants have adaptations that allow them to thrive on land —>
structural adaptations to gain resources from air and soil
Plants have adaptations that allow them to thrive on land
—> structural adaptations to gain resources from air and soil
Shoot system and root system
pollen and egg
Plants have adaptations that allow them to thrive on land
—> structural adaptations to gain resources from air and soil
Shoot system:
Photosynthetic structures exposed to light and air aboveground
Plants have adaptations that allow them to thrive on land
—> structural adaptations to gain resources from air and soil
root system:
Belowground anchorage structures exposed to
water and minerals in soil
Plants have adaptations that allow them to thrive on land
—> structural adaptations to gain resources from air and soil
Pollen:
Protects sperm, allowing them to be carried by wind or animals
Plants have adaptations that allow them to thrive on land
—> structural adaptations to gain resources from air and soil
Egg:
Remains in female
reproductive organ, allowing the embryo to develop within the mother plant’s body
Plant structure
Vascular System:
Network of tubes that transports water, sugar, and minerals
Plant structure
Cuticle:
Waxy coating that prevents water loss
Plant structure
Leaves:
Main site of photosynthesis and gas exchange through
Plant structure
Lignin:
Chemical in cell walls that stiffens them, increasing suppor
The evolutionary history of plants records adaptations for
living on land
The evolutionary history of plants records adaptations for living on land
There are four major groups of modern plants:
Bryophytes
Seedless Vascular plant
Gymnosperms
Angiosperms
Major events in the evolution of plants highlight adaptations for
dry land
Major events in the evolution of plants highlight adaptations for dry land
Bryophytes
(mosses, liverworts, hornworts, 475 mya): evolution of waxy outer layer reduced water loss; still dependent on water for reproduction.
Major events in the evolution of plants highlight adaptations for dry land
Ancestral Green algae
origin of first terrestrial adaptions (bout 475 mya)
475 million years ago: nonvascular plant
Origin of Vascular tissue (about 425mya)
origin of seed (about 360 mya)
Bryophytes are seedless, nonvascular plants
Bryophytes have adaptations for land:
Cuticle,:
a waxy outer layer which retains moisture
§Structures that provide internal protection for gametes and embryos
Bryophytes are seedless, nonvascular plants
§They lack
seeds and vascular tissue.
Bryophytes are seedless, nonvascular plants
Modern bryophytes can survive
only in watery environments
Bryophytes are seedless, nonvascular plants
Mosses are
the most common bryophytes.
Three Types of Bryophytes
§Mosses (most common)
Liverworts
Hornworts
Moss diversity
Red moss capsules
Peat moss bog (3% of Earth’s land surface)
Moss colony on a tree in a coastal forest
Bryophytes come in two distinct forms representing an alternation of generations
•Diploid Sporophyte
produces spores (haploid reproductive cells) which are tough and can survive harsh conditions.
Bryophytes come in two distinct forms representing an alternation of generations
•Unlike most plants, in bryophytes the haploid gametophyte is
dominant (larger & more obvious).
Bryophytes come in two distinct forms representing an alternation of generations
Parts:
Spore Capsule
Sporophytes
Gametophyte: green, sponge-like, forms a mat
Brachiopoda
§Lamp shells
§Bilateral symmetry
§Lophophore
§Similar to bivalves
§Suspension Feeders
Cephalopoda
§Molluscs
§Bilateral symmetry
§Large heads
§Foot modified into tentacles
§Shelled or unshelled
§Relatively smart
§Active predators
The first vertebrates with jaws were
cartilaginous fishes
The first vertebrates with jaws were cartilaginous fishes
Cartilaginous fishes
(sharks and rays, ~1,000 species) have flexible skeletons made of cartilage and use two pairs of fins and a tail to swim, fast agile swimmers.
-First vertebrates with jaws
The first vertebrates with jaws were
cartilaginous fishes
The first vertebrates with jaws were cartilaginous fishes
Evolution of jaws, along with a stomach
Shift from filter feeding and scavenging à active predation
Most chondrichthyans (cartilaginous fishes) are predators.
No sharks yet
Silurian
443.8-419.2mya
Vascular plants
Jaws and bones in fish
Warmer, but many climate fluctuations
Vascular tissue was an important
evolutionary adaptation
Vascular tissue was an important evolutionary adaptation
§Nearly all plants (except for bryophytes) have
vascular tissue.
Vascular tissue was an important evolutionary adaptation
§Nearly all plants (except for bryophytes) have vascular tissue.
these would be
SEEDLESS VASCULAR PLANTS
GYMNOSPERMS
ANGIOSPERMS
Phloem primarily transports
sugars
Phloem primarily transports sugars
Phloem
contains living cells that transport sugars from where they are made or stored to other parts
of the plants.
Phloem primarily transports sugars
Phloem sap always flows from a
sugar source to a sugar sink.
Xylem primarily transports
water and minerals from the roots
Xylem primarily transports water and minerals from the roots
Xylem
made of dead cells acting as tiny pipes.
Xylem primarily transports water and minerals from the roots
§Xylem sap is moved through the plant body via
transpiration.
Xylem primarily transports water and minerals from the roots
Xylem sap is moved through the plant body via transpiration.
Water evaporates into air from the leaves —>
pulling up more water from the roots, through root hairs.
Xylem primarily transports water and minerals from the roots
Xylem sap is moved through the plant body via transpiration.
Within the xylem, water molecules stick to
the interior cell walls and each other (hydrogen bonds)
Ferns are an example of a type of
seedless vascular plant
Ferns are an example of a type of seedless vascular plant
§Seedless vascular plants (ferns) have important adaptations for living on land:
§Vascular tissue
§Lignin-hardened cell walls
Ferns are an example of a type of seedless vascular plant
Lycophytes
1st appeared 410 mya, long surviving group
Ferns are an example of a type of seedless vascular plant
Carboniferous forest:
forest of seedless vascular plants of 360-300 mya
Ferns are an example of a type of seedless vascular plant
Modern ferns:
: found in tropics and temperate woodlands
Fern Diversity
Tree fern
Staghorn fern
Young fern ‘fiddlehead”
Foxtail fern
Anatomy of a Fern
Frond
•large leaves with many leaflets
Anatomy of a Fern
Lignin
•stiffen cells and allows the plant to stand up tall and strait
Anatomy of a Fern
Spore Capsule:
•: structures on underside of leaves, containing spores
Chelicerata are
a group of eight-legged arthropods
Chelicerata are a group of eight-legged arthropods
§Most are
terrestrial carnivores.
Chelicerata are a group of eight-legged arthropods
§Early forms like xiphosura
(horseshoe crabs) evolved during the Silurian
Chelicerata are a group of eight-legged arthropods
examples:
Yellow desert scorpion
Wood tick
Dust mite
Horseshoe crab
Mollusk Groups
examples:
Gastropods- (snails and slugs) Blackmargin sea slug
Bivalves -(clams, mussels, and oysters) Bay scallop
Cephalopods -(squid and octopi)European squid
Gastropoda
Marine and Terrestial
Hermaphrodites
Torsion
Muscular foot
Bivalvia
2 valves to the shell
Filter feeders
A Gallery of Mollusks
Lampsilis mussel, Chambered nautilus, Invasive apple snail, Blue ringed octopus
Sacoglossa
“Sap sucking”
Solar Powered!
There are more species of bony fishes than any other group of vertebrates
species ammout
28,000
There are more species of bony fishes than any other group of vertebrates
Bony fishes
first group to have internal skeletons reinforced with calcium.
They also have a swim bladder to maintain buoyancy, and an operculum that circulates water over the gills.
A Gallery of Bony Fishes
Flying gurnard, Thorny seahorse, Yellow-ribbon sweetlips, Chevron barracuda
The lobe-finned fishes are an evolutionary branch of the
bony fishes
The lobe-finned fishes are an evolutionary branch of the bony fishes
Lobe-finned fishes
have muscular fins supported by rod-shaped bones that are homologous to amphibian limb bones.
The lobe-finned fishes are an evolutionary branch of the bony fishes
Include the
lungfishes and coelacanths.
Devonian
§Relatively warm
§High sea levels
§Ended with an extinction
§Plants spread further into the
The gymnosperms were
the first plants to evolve seeds contained within cones
The gymnosperms were the first plants to evolve seeds contained within cones
A seed consists of
an embryo and food supply inside a protective coating.
The gymnosperms were the first plants to evolve seeds contained within cones
Seeds are one of the key adaptations that allowed
plants to spread across the land.
A Gallery of Gymnosperms
Conifers: most abundant; pine, spruce, cedar, fir, and redwood
Bristlecone pines: live up to 4,500 year, native to California
Cycads: diverse group found in the tropics
Redwoods: largest and tallest trees in the world. These trees can live for thousands of years.
Ginkgo trees: found in fossils dating back 270 million years, native to China
Gymnosperm Reproduction
Cones of most gymnosperms house
all reproductive stages
Spores, eggs, sperm, zygotes, and embryos
Gymnosperm Reproduction
Pollen-producing cone —>
develop male gametophyte (haploid sperm) within pollen grains produced from spores
Gymnosperm Reproduction
Ovule-producing cone —>
develop into female gametophyte (haploid egg)
Gymnosperm Reproduction
Fertilized egg (diploid zygote)
develops into a seed
Embryos are provided a food supply and protective coating within the seed
Cones of most gymnosperms house
all reproductive stages
_______, by far, outnumber all other animals combined
Insects
Insects, by far, outnumber all other animals combined
Most have easily recognizable segments (head, thorax, and abdomen), three pairs of legs, and one pair of antennae.
Consumers or detritovores, some parasites
Huge diversity
Mostly terrestrial
Many metamorphose
Insect Diversity
examples:
Hemiptera (true bugs), Hymenoptera, Coeloptera, Lepidoptera,
Arachnida
Eight-legged arthropods
Head
Abdomen
Book lungs
Many predators
Some parasites
Millipedes and centipedes are easily recognizable
Millipedes and centipede have
§segmented bodies, which appear as many fused rings, each with jointed appendages attached.
Millipedes and centipedes are easily recognizable
Mili-multiple per segment
Centi-1 per segment
All are land dwelling
Sharks
Ancestors to modern sharks
Dorsal fins
Unable to replace teeth
Amphibians
have four limbs and were the first to occupy land
Amphibians have four limbs and were the first to occupy land
All terrestrial vertebrates are tetrapods,
animals with four limbs
Amphibians have four limbs and were the first to occupy land
Amphibians
were the first tetrapods
Amphibians have four limbs and were the first to occupy land
tetrapods and Amphibians were followed by the evolution of
amniotes (reptiles & mammals).
Amphibians live on land, but must
reproduce in water
Amphibians live on land, but must reproduce in water
Biphasic lifestyle —>
blend of aquatic and terrestrial adaptations.
Amphibians live on land, but must reproduce in water
Aquatic adaptations:
§Typically breed in or near water
§Eggs lack shells (like in fishes)
§Larval stage has gills, lateral line
Amphibians live on land, but must reproduce in water
Terrestrial adaptations:
Adult has lungs
Musculoskeletal system with four limbs
Moist skin
A Gallery of Amphibians
7,089 species of frogs & toads
Frogs- have powerful hind legs that allow hopping.
Caecilians -superficially resemble earthworms or snakes. (~212 species)
Toads-live entirely on land.
Salamanders-can be either aquatic or terrestrial. (~731 species)
Early amphibians
370mya
Fins to limbs
Mississippian and Pennsylvannian
358.9-298.9mya
”Carboniferous”
Mass expansion of land by plants
Winged Insects
Reptiles
Reptiles, including birds, can complete their entire life cycle
on land
Reptiles, including birds, can complete their entire life cycle on land
Reptiles have two major adaptations for land:
.They reproduce via an amniotic egg.
−Hard, waterproof shell
−Provides an aquatic habitat for the embryo
Reptiles, including birds, can complete their entire life cycle on land
Reptiles have two major adaptations for land:
2.
Reptiles also have waterproof skin.
Also they are ectotherms (more on this later)
Turtles have hard shells
~356 species
All turtles have protective shells made from to shields fused to vertebrae
Aquatic and terrestrial
largest living reptiles
Crocodilians