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organ system
shoot system: vegetative organs (leaves & stems), reproductive organs (flowers and fruits)
root system
determinate growth
maximum size is genetically determined (usually cannot heal/regrow)
scientists
robert hooke: observes cells
schleidern & swann: cell theory
virchow: cells come from cells
all cells contain
plasma membrane, cytoplasm, dna, ribosomes
ribosomes
non-organelles that synthesize proteins
ER
modify proteins & synthesize lipids
features unique to plant cells
cell wall, central vacuole, plastids (chloroplasts)
tonoplast
membrane of central vacuole. plasmolyzed, flaccid, or turgid
label chloroplast
pigments stores in thylakoids

middle lamella
thin layer of pectin binding adjacent cells
primary cell wall
occurs on surface of all plant cells. composed of microfibrils that are thin and elastic, allowing for cell enlargement
secondary cell wall
made of layers deposited inner to primary wall, occurs when cells reach mature size. lignin + cellulose = rigidity
cell wall expansion
longitudinally but not laterally (up and down like a spring)
plasmodesmata
pores that allow for communication between cells
all tissues originate from
meristematic tissue. totipotent: can regenerate entire plant
meristems
regions of continuous cell division and growth
3 types: apical meristems, lateral meristems, intercalary meristems
apical meristem
located at very end of stems & roots
produce 3 types of primary meristematic tissues: protoderm, ground meristem, procambium
protoderm
gives rise to epidermis
ground meristem
gives rise to ground tissue (photosynthetic, storage, support)
procambium
gives rise to vascular tissue (transport)
lateral/secondary meristem
responsible for secondary growth (increases girth)
-vascular cambium (from procambium): gives rise to vascular tissue
-cork cambium (from pericycle & cortex): produces peristem
seeds allow populations to
-propagate next gen
-maintain genetic diversity
-survive harsh conditions (dormancy)
-disperse
typical seed contains
-embryo 2n
-nutrient source (endosperm 3n and/or cotelydon)
-seed coat
embryogenesis in eudicots
developing seed contains endosperm & zygote
zygote forms proembryo anchored by suspensor
embryo proper in globular stage
heart stage (cotyledons)
torpedo stage (shoot apical meristem)
mature embryo: radicle, hypocotyl, epicotyl, seed coat
seed coat components
testa: outer coat
tegmen: inner coat
hilum: scar where seed attached to endocarp (fruit)
micropyle: where pollen tube entered
plume
young shoot containing shoot apical meristtem & leaf primordia
radicle
embryonic root
hypocotyl
region between cotyledon attachment point and radicle
epicotyl
region between cotyledon attachment point and shoot tip
monocot seeds
pericarp is fused with seed coat, testa and tegmen fused
root tip protected by coleorhiza, plume by coleoptile
scutellum
single cotyledon in monocots
starchy endosperm
aleurone: outer layer, secretes enzymes during germination
dormancy barriers in seed coat
-physical dormancy: impermeability to water or oxygen
-chemical dormancy: compounds in seed coat inhibit germination
dormancy barriers in embryo
-physiological dormancy:embryo requires environmental conditions to complete maturation
-endodormancy: internal biochemical processes must be met before germination
-ecodormancy: external factors are not optimal
steps required to initiate germination
imbibition
enzyme activation
breakdown of starch into sugar
types of germination
epigeous: hypocotyl (plume hook) eloongates, cotyledons extend above ground
hypogeous: epicotyl elongates, cotyledons remain belowground
what emerges first
radicle
epigeous monocots
monocot bends into hook and emerges before coleoptile
area of cell elongation
right behind apical meristem, below area of maturation
plant anatomy vs morphology
morphology studies physocal form & external structure
anatomy studies the tissues and cell structures of organs
rhizosphere
thin area around roots where complex chemistry occurs for nutrient absorption
site of gravity perception
root cap
pericycle
outer boundaru of vascular cylinder, zone of root branching
epidermis
single layer of cells at outer boundary, arises from protodern. might elongate into sticky root hairs
cortex
several layers between epidermis and vascular cylinder, differentiates from ground meristem.
endodermis: innermost layer, helps control mineral accumulation
casparian strip: waxy material embedded in walls, forces movement through cells
vascukar cylinderr
vascular tissue that differentiates from procambium, occupies center of root.
consists of primary xylem, primary phloem, and peicycle.
in dicots, forms a central core in 2+ points, phloem in between radial branches of xylem. in monocots, central pith surrounded by rings of xylem & phloem
xylem tissue
tracheids and vessel elements
protoxylem: first xylem element to mature, capable of transporting water while elongating (form annural rings)
metaxylem: mature after cell elongation is completed. form thick secondary walls for water & mineral transport. crush protoxylem cells as they grow
phloem tissue
forms sieve tubes with companion cells
first part of vascular system to become functional
vascular cambium
in long-lived dicota, parts of pericycle cells combine wiith residual procambium
secondary phloem
forms inner bark, pushes pericycle outwards. remaining pericycle becomes cork cambium
bark & wood
bark: secondary phloem
wood: secondary xylem
true leaves arise from
leaf primordia
mesophyll
middle layer of cells in leaf
includes palisade and spongy tissue
cuticle
waxy layer that prevents water loss
guard cells
surround stomata, supported by subsidiary cells
when full of water, stretched open
trychomes
hairs that can secrete substances
makes leaf pubescent (hairy) instead of glabrous (shiny)
palisade cells
upper part of mesophyll, contains chloroplasts and are main tissues for photosynthesis
spongy cells
lower part of mesophyll, main function is to allow diffussion of gasses
veins
vascular bundles that form a network of xylem & phloem
stem tuber
modified stem used to store food underground
bulb
short, flattened stem-bearing fleshy, food-storing leaves
rhizome
underground, horizontal stem
shoot system has origin in
plume of the embryo
stem functions
support, conduction, photosynthesis, food storage
nodes
attachment points for leaves
axillary bud
area between a leaf and the stem, can give rise to new axillary shoot
types of meristematic tissue
-protoderm (epidermis)
-ground meristem (ground tissue)
-procambium (vascular tissue)
secondary growth
increase in thickness or girth of the stem caused by cell division of secondary meristem
consists of vascular cambium & cork cambium→ periderm
vascular cambium divides into
secondary xylem on inside and secondary phloem outside
periderm
replaces epidermis in shoots at end of first year growth. mostly impermeable due to waxy suberin
new layers produced inside the older layers (rhytidome), which die and are pushed out
bark
all tissue external to vascular cambium
inner bark: secoondary phloem, cortex, phelloderm
outer bark: cork cambium, periderm, cork, etc.
wood
consists of original pith, primary xylem, and secondary xylem
phloem on outside, xylem inside
heartwood
oldest (innermost) layers of secondary xylem which no longer conduct water
tylosesvessel prevent waterflow
sapwood
water-conducting xylem in periphery of heartwood
bundle scars
circular markings where leaves fall off
types of branching
monopodial: buds do not degrade, and all shoots continue to grow
sympodial: terminal buds degrade, and lateral buds closest to the apex become the new terminal shoot
true leaves
never in the apex, aways substend an axillary bud
petiole
stem-like part of leaf, connects blade to stem
sessile
leaves without visible petiole
peltate
petiole attaches underneath blade
perfoliate
stems pass through center of blade
phyllotaxy
arrangement of leaves on stem
reticulate venation
netlike pattern

palmately lobate
lobes outline palmate venation
toothed leaves
serrate: sharp teeth pointing forward
dentate: sharp, symmetrical teeth
crennate: teeth are rounded (mini bumps)
ovate
widest toward base
obovate
widest toward apex (top)
elliptic
single wide middle position
oblong
longer wide middele portion
attenuate
tapering to a point
acuminate
curving inward to a point
cuneate
wedge-shaped base
cordate
upside-down heart (base)
primary growth only
herbaceous stems
herbaceous stem anatomy
epidermis, cortex, pith, primary xylem tissue & primary phloem tissue, cambium layer

glabrous vs pubescent
glabrous: without hairs, pubescent: hairy
vascular bundles, derivatives from
xylem and phloem are arranged in bundles that arise from procambium bundles
xylem arises from cell facing inside (conducts from roots to rest of plant)
phloem arises from cell facing outside (distributes food from photosynthetic tissues to other tissues)
vascular bundle arrangement
solenostele: some eudictos, tissues in continuous ring
eustele: most eudicots. vascular bundles discrete but in a circle
atactostele: most monocots, vascular bundles scattered through stem
phloem tissue is composed of
sieve cells and companion cells
sieve cells
aranged in tube columns, end walls called sieve plates.
living cytoplasm but no nuclei. companion cells possess nuclei and perform all processes to keep them alive