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Genotype frequency
(Number of individuals that have a genotype) / (total number of individuals in the population )
Allele frequency
(number of copies of an allele) / (Total number of of alleles in a population)
Evolution
Change in the genetic makeup of a population over generations
Natural Selection
“Variants within populations that can be inherited and that are best suited for growth and reproduction in given environment
Environmental Variants
(sub category from natural selection) due to differences in the environment
Genetic Variations
(Sub category from natural selection) Due to differences in the genetic material that is transmitted from parent to offspring
Mutations
Arise either from random errors during DNA replication or from environment factors that can damage DNA
Evolution Theory
Understanding of how evolution works based on many observations and experiments
Example of evolution theory
Closely related species are likely to resemble each other more closely than they do more distantly related species
Phenotype variation
Differences in observable traits among individuals of our species
Species
group of individuals that are capable, through reproduction, of sharing alleles with one another
Gene pool
Consists of al the alleles present in all of the individuals in the species
Population Genetics
Study of genetic variation in natural populations
Somatic mutations
Occurs in bodies tissues in non-reproductive cell; only affects only the cell descended from the one cell in which the mutation originally arose - not passed on to the next generation
Germ-line mutation
Present in sperm or egg, are evolutionary important because they are passed on to the next generation and appear in every cell of an offspring
how to determine evolution has occured
If allele frequencies change or if genotype frequencies change over a generation
Hardy-Weinberg equilibrium
Describes what we expect at a gene (locus) when evolution is NOT occuring - baseline expectation for a non-evolving population
5 things that are said to be Hardy - Weinberg equilibrium as these are meat
No selection: no differences in the survival and reproductive success
No mutations: No mutations within gene
No migration: Population size is not changed by movement of individuals
Large population ( No genetic drift): Sufficiently large to prevent sampling errors
Random mating: Individuals in the population mate at random
Natural Selection occurs when these observations are met
There is variation in the population
When the variation is heritable
resources must be finite driving competition among individuals
As a result of competition, mortality occurs
evolution by natural selection
Leaving more offsprings with a certain trait results in the trait increasing in frequency in the next generation
Fitness by natural selection
differential survival and reproduction of individuals with certain traits in the given environment
Fitness
Measures of ability of an individual to survive and reproduce in a particular environment, leaves its genes in the enxt generation
Adaptations
Traits that increase fitness; new genes to survive new environment
Positive Selection
Natural selection for favorable alleles → increases the frequncies of a favorable allele
Negative Selection
Natural selection against harmful alleles → decrease allele frequency of a harmful allele
Balancing Selection
Selection favoring allelic variation → Multiple alleles are maintained at intermediate levels
Stabilizing selection
Results in selection against extreme phenotypes
Directional Selection
Only selects for or against extreme phenotype; shifts the distribution of a phenotype so the mean value increases or decreases in the population ( Graph shifts left or right )
Disruptive Selection
results in selection against intermediate phenotypes and is thought ot be important for certain speciation processes
Artificial Selection
[ Form of directional selection ], successful genotypes are selected y the breeders, not by competition
Indirect Competition
One sex chooses members of the other sex based on phenotype criteria
Direct Competition
Between individuals of the same sex
Inter selection
Focuses on interactions between male and female ( Indirect Competition )
Intra selection
Males fighting for access to females ( Direct Competition )
Adaptive mechanisms of evoluton
Driven by environmental pressures
Favors traits that improves survival and reproduction
Leads to better fit between organisms and environment
ex. Direction, disruptive, stabilizing, balancing, and sexual selection
Natural Selection
Non-Adaptive Mechanisms of Evolution
Not driven by natural selection or fitness differences
Occurs regardless of whether traits are beneficial, harmful, or neutral
Can increase or decrease genetic variation by chance
ex. genetic drift, genetic flow (migration), mutations
Genetic Drift
[non-adaptive], allele frequencies change due to chance - depends on population size and allele change over generations
Population bottleneck
A population is drastically reduced in size - negative results in environment resulting in love survival rates
Founders effect
occurs randomly, reduces genetic diversity - Occurs when a few individuals become isolated from a larger population
Migration
Connects population by allowing gene flow between them, resulting in allele frequencies to change
Gene flow
population becomes more genetically similar over time - matting with individuals of another population and becoming all one
What must occure for a popualtion to be consisdered seperate species
Population results be reproductively isolated
Pre-Zygotic
Reproductive isolation happens before fertilization
Post-Zygotic
Reproductive isolation happens after fertilization
Pre-Zygotic Mechanisms
Geographical / Ecological
Temporal
Behavioral
Gametic
Mechanical
Pre-zygotic Geographical / ecological
Individuals are separated in space or habitat
Pre-zygotic Temporal
Plant flowers at different times, and animals are active at different times
Pre-zygotic Behavioral
Individuals only male with other individuals based on specific behavior
Pre-zygotic Gametic
Incompatibilities exist between gametes
Pre-zygotic Mechanical
Species reproductive organs are incompatible
Post-zygotic hybrid inviability
After fertilization, a zygote fails to develop
Post-zygotic hybrid sterile
Offspring are produced but are sterile
Speciation
the process of one species splitting into and species as a by product of genetic divergence
Allopatric Speciation
Barrier forming between both species
Initial speciation
Can result in evolution during each population diverging until they are reproductively isolated
Dispersal (Allopatric)
Individuals colonize a new area
Vicariance (allopatric)
Where a geographic barrier arises, a population splits into two separate population - overtime, the time population will become genetically distinct from one another until speciation occurs
Disruptive Speciaiton
Can drive reproductive isolation in the absence of physical separation
Sympatric Speciation
Can be driven by hybridization - in hybridization, the number of chromosomes can be different from either parental species → as this occurs, hybrids are reproductively isolated from parents because of gamete incompatibilities
Sister taxa
Groups that an immediate common ancestors not shared by any other group
Phylogentic Trees
Shows pattern of descent, does not generally show how old lineage are or how much they have changed
Example of Stabilizing Selection
In a fish species, the eggs tend to be fertilized more often by the mid-sized males and less often by the largest and smallest males - results in selection against extreme phenotype
Example of directional selection
Environmental conditions - food source characteristics - causes selection towards one extreme of a traits range of variation
Polyphyletic
Consists of taxa that share a trait that is not present in their most recent common ancestor
Paraphyletic
Incudes a common ancestor and some but not all of the descendants
Monophyletic
Includes a common ancestor and all of its descendants
Recombinations and Mutations
Produce new alleles in a population
Genetic Variation
Critical for evolution because it is the source material for natural selection