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Plant Responses
cannot get up and move away from a predator or poor environment
If a resource becomes available, plants have to grow towards it
Two ways plants get larger
Cell division and Cell elongation
All growth is regulated by
hormones
Auxins
stimulate elongation, division, and differentiation
Promote elongation of cells in
coleoptiles and stems
Heavily concentrated in (Auxins)
Shoot and root apex, Expanding leaves and buds, and Developing seeds and fruits
Auxin Effects on Whole Plant
influence growth, cell differentiation, and responses to light and gravity
Apical dominance driven by
auxin production
If apical shoot has poor dominance,
Then the plant can have “poor” form
Polar transport
from tip to base of auxin within the plant
Gibberellins stimulate
elongation
High concentrations in (Gibberellins)
Stem and root apex, Buds, embryos, cotyledons, Flowers and fruits, Vascular cambium
Promote stem elongation help seeds and buds
break dormancy
Effects of Gibberellins
include promoting stem elongation, breaking seed dormancy, and influencing flowering
Cytokinins
stimulate division
Inhibits elongation
Synthesized mostly in (Cytokinins)
root tips, also in embryos, young fruits, young leaves, root tips
Stimulate cell division and retard leaf aging
Can stimulate auxin production, Inhibits elongation, but stimulates enlargement
Ratios between cytokinins and auxins will determine the
differentiation of tissue within the plant
Ethylene
gas stimulates aging, senescence
Ethylene promotes
fruit ripening, senescence and abscission
Modifies effects of auxin on growing organs
Inhibits stem and root elongation
Abscisic acid
is a general inhibitor of growth
Abscisic Acid Counteracts auxins and gibberellins
at higher concentrations it promotes seed dormancy and stress responses
Promotes stomatal closure (Abscisic Acid)
May trigger seed and bud dormancy and also promotes abscission
Roots experiencing drought produce abscisic acid and load into xylem (Abscisic Acid)
Signals stomata in leaves to close, reduce transpiration
All growth responses to the environment rely on
hormone production
Some environmental responses/movements are not
related to growth or hormones
Tropisms
are directional growth related to an external stimulus
Nastic movements
are non-directional and not related to an external stimulus
Phototropisms
Responses to light
Growth responses to directional light source
Blue light receptors trigger auxin transport
Auxin triggers differential cell elongation
Gravitropism
Orients plant parts to the pull of gravity
Mechanical stimulus (Gravitropism)
from amyloplasts (statoliths)
Triggers auxin transport
Differential elongation
Stems (Gravitropism)
negative gravitropism as they grow upwards away from gravity.
Roots (Gravitropism)
positive gravitropism as they grow downwards towards gravity.
Thigmotropism
Responses to physical contact
Usually see it with stems or tendrils
Cells in contact with the object shorten, opposite cells elongate - Not clear if auxin gradients are involved
how roots navigate around rocks in soil
Thigmotropism
Photoperiodism
Responses to change in relative length of daylight and darkness
Photoperiodism also allows
biological clock to be “reset” for seasonal adjustments to day length
Photoperiodism is controlled by
phytochrome pigment
Phytochrome
Converts to active form (Pfr) during daylight, or when exposed to red wavelengths and helps plants detect changes in light duration
Phytochrome physically interacts with
phytochromeinteracting factors (PIFs) to regulate gene expression in response to light conditions
Nastic Movements
Reversible responses to nondirectional stimuli
Movements in leaves, leaflets, flowers