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heterogeneous
patches of less suitable habitat in the mosaic require adaptations to maximize fitness
habitat fragmentation
rapid alteration of landscape spatial structure including deleterious genetic, population, and species effects on resident species
research as focused on _________ ______________ rather than ________________ _____________ that minimize biodiversity loss
patch dynamics (conservation approach), land-use methods (sustainable production approach)
important to consider ___________ of habitat remaining and ______________ _______________________ of habitat patches
amount, spatial configuration
habitat loss is a direct result of
fragmentation
island biogeography theory
explains how species come to be distributed among oceanic islands
fragmentation usually results in declining ___________ ___________ through time
patch area:
- results in many small patches
population level
limited resource availability, reduced colonization rates, altered reproductive success
limited potential population size from:
1. environmental stochasticity (all individuals)
2. demographic stochasticity (some individuals, small N)
3. natural catastrophes
4. reduced genetic diversity (genetic drift)
5. altered resource availability and the interaction with 1-4
community level
changes in patch species composition
- dependent on species requirements and dispersal abilities
- evolution of metapopulation and metacommunity ecology
- reduction of richness in small patches
fragmentation increases edge effects
smaller patches --> relatively greater edge habitat
- patch size reductions increasingly expose interior forces
- edges hotter, drier, windier, higher wavelength intensities, composition --> altered animal composition
- edge effects inclde edge extent and magnitude
fragmentation results in various patch shape complexities
edge:area ratio useful metric, minimized with perfect circle
greater shape complexity
1. increase colonization rates
2. increase environmental heterogeneity
3. reduce core area for interior patch specialist
4. research indicates:
a) shape complexity increases exponentially with increasing patch size
fragmentation results in increased patch isolation
- changing characteristic in time and space, depends on:
1. time since isolation
2. distance between patches
3. temporal changes in matrix characteristics
4. dispersal traits of species
species relaxation
initial richness declines through time as species cannot maintain viable populations
habitat corridors
1. allow movement between isolated populations
2. potential for both positive (dispersal, genetic connectivity) and negative (spread of diseases, invasive species) impacts
fragmentation effects related to matrix quality
1. depends on organisms involved
2. higher (organic farm, high shade) and lower quality (traditional farm, low shade) matrix environments
fragmentation effects in natural systems factors:
1. interactions between drivers of fragmentation
2. poor understanding of species life histories
3. temporal changes in fragmentation effects
4. synergistic interactions between fragmentation and other processes
habitat degradation can be viewed as alteration of the ________________ and _________________ environments
biophysical alteration: invasive species, species irruptions, altered hydrology
chemical alteration: nutrient increases/decreases, herbicides/pesticides, industrial pollutants, acid deposition
marine habitats biophysical and chemical impacts
biophysical: climate change, altered currents, upwellings, sea level rise, reef loss
chemical: nutrients --> dead zones, acidification --> massive biodiversity loss