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Aristotle
-Fixed ideal species
-Scala Naturae (scale/ladder of nature) Humans at the top, followed by mammals, birds, etc.
Age of Reason (1700s-1800s)
Birth of science. (start of observations, thinking, and reasoning)
Linnaeus
-created the orderly binomial naming system
James Hutton
Father of gradualism (earth is always changing geographically over time)
Charles Lyell
-Father of geology
-Uniformitarisnism: same geographical processes in past as today
-EARTH IS EXTREMELY OLD
Erasmus Darwin
Natural philosopher, Development of life over eons
Lamark
-Naturalist
-Extinct species have been replaced by descendants w/ new features
-First to propose mechanism for evolution
-Lamarkism: inheritance of acquired traits (WRONG)
Charles Darwin
-Naturalist
-Collected a lot of wildlife, plants, and fossils from the Galapagos islands
-Famous for the theory of evolution
The Theory of Evolution
-Darwin wrote his essay in 1844
-Alfred Wallace came up with same idea and wrote Darwin in 1858
-Both co-published in 1858
Origin of Species (1859)
1. Descent with Modification
2. Natural Selection
Natural Selection
-unique idea by Darwin
-"descent through modification"
-Natural Selection is a mechanism for evolution
Natural Selection & evolution
1) heritable variation
2) can enhance small variations through artificial selection
3) All species produce more offspring than the environment can hold
4) Adaptations that help survivability can accumulate for max. success
5) Environment controls evolution
NOT explained in Darwin's Theory
origin of life, how variation arises, how inheritance works, why variation still exists, "sudden" changes in fossil record, Source of totally "new" characters (Most are now explained)
Evidence of Darwin's Theory
1. Fossil record
2. Homology
3. Convergence
4. Biogeography
5. Molecular Biology
Homology
-similarity in structure from common ancestor
-structure modified for different functions
analogous structures
-Body parts that share a common function, but not structure
-different species
Vestigial Structures
-Homologous
-no present adaptive function
Convergence
-analogous
-natural selection acted the same in same conditions
Biogeography
-distribution of species via geographic history
Gene Flow
Movement of alleles across a species' range
Ways to define species
Morphological species
Ecological Species
Phylogenetic species
BIOLOGICAL SPECIES
Biological species concept
Organisms that are reproductively isolated from each other are different species. biological barrier
Morphological Species Concept
Organisms that have significant morphological and anatomical differences are different species
Pre-zyotic Barriers
Prevent fertilization: Temporal isolation (ex: different mating times), Behavioral isolation (ex: different mating calls/courtship cues, Habitat isolation (ex: ecological, different habitats), Gametic isolation (ex: gametes cant fuse-different flowers have different pollen), Mechanical isolation (ex: certain dog breeds cannot breed with each other bc of size)
post zygotic barriers
act after a hybrid zygote is formed, reduced hybrid viability (may be born but fails to develop), reduced hybrid fertility (born but sterile),hybrid breakdown (1st generation is vaible but 2nd generation fails)
Gene & Reproductive isolation
A gene may influence reproductive success, snails can only mate with other snails with the same shell rotation
Problem with biological species concept*
-fossil species
-asexual species
-hybrids do happen
Speciation
Formation of new species
Allopatroc Speciation
-In which a population forms new species because its geographically isolated from the parent population (Panama land bridge, mosquitofish)
adaptive radiation
An evolutionary pattern in which many species evolve from a single ancestral species (finches), organisms diversify due to environmental changes ex: Hawaii Honeycreeper bird -> evolved to multiple other birds with different characteristics
What causes speciation?
changes (mutations), sexual selection, reproductive incompatibility
sympatric speciation
The formation of new species in populations that live in the same geographic area (cichlids and muddy water), Same geographical area, rare
Chromosomal Errors
-sympatric speciation
-aneuploidy 2n + 1 or 2n - 1 (too many or too few chromosomes, non disjunction)
-polyploidy 4n
habitat differentiation
sympatric speciation can also result from the appearance of new ecological niches
Gradual speciation
species diverge gradually over time in small steps
punctuated equilibrium
species evolve during short periods of rapid change followed by long periods of stasis
allopolyploidy
-polyploidy resulting from contribution of chromosomes from two or more species (rarely found in animals, mostly plants)
-normal gamete + polyploid gamete = polyploid gamete -> second mating -> allopolyploidy
Phenotypic relations
mostly genetic but environment can influence expression
Discrete genetic variation
Variation in a single gene locus
Continuous Variation
phenotypes produced by combined effects of 2 or more genes
How do new alleles arise?
mutations in the DNA
What is a population?
a group of individuals of the same species living in the same area at the same time
Allelic frequency
the percentage of any specific allele in the gene pool
Genetic frequency
refers to the percentage of each allele for a particular trait in a population
Microevolution
any change in a populations allelic genotypic frequency over time
Hardy-Weinberg Principle
principle that allele frequencies in a population will remain constant unless one or more factors cause the frequencies to change
Hardy-Weinberg conditions
1. No mutations
2. Random mating
3. No natural selection
4. Extremely large population size
5. No gene flow
Hardy-Weinberg equation
p2 + 2pq + q2 = 1
What if a population meets the H-W equation?
every generation over time will have the same genotypic rations
What is H-W used for?
-Used to determine population's genetic makeup
-if the actual ratios do not equal HW ratios then the population is evolving
Mechanism of microevolution
1. natural selection
2. genetic drift
3. Founder Affect
4. Bottleneck Effect
5. Gene Flow
Natural Selection
-acts non-randomly on phenotypes of individuals
-changes allelic & genotypic frequencies of populations
-always leads to adaptation of population to current environment
Aspects of natural selection
1) relative fitness
2) directional selection
3) disrupted selection
4) Stabilizing selection
Genetic Drift
-random
-occurs in small populations (like sampling errors in statistics)
Outcomes
-one allele may become fixed
-random changes in allele frequency
Founder Affect
-genetic drift resulting from the colozination of a new isolated population
-founder gene pool is different from original species
-small population —> more drift
-superior alleles may be lost
Bottleneck Effect
-genetic drift that dramatically cuts population size
-gene pool survivors are lost
Gene Flow
-alleles move in/out of population
-migration
-dispersal of gametes, seeds, larvae
Results of gene flow *
-adds diversity to population
-reduce differences between populations
Forms of Natural Selection
-Directional
-Diversifying
-Stabilizing
-Frequency-Dependent Selection
-Sexual Selection
Macroevolution
Gene flow + 3.8 billion years
Relative Fitness
Best reproductive success
Directional Selection
-shifts populations genetic variance toward the new, fit phenotype
-selects phenotypes at one end of the spectrum (mouse fur color)
Diversifying Selection
-intermediates are less fit than extremes
-maintains diversity
-increases genetic variance
Stabilizing Selection
-intermediate types more fit than extremes
-decreases genetic variance
Frequency-Dependent Selection
-the fitness of a phenotype depends on how common
Sexual Selection
=success based on traits related to obtaining mates
-leads to sexual dimorphism
Intrasexual Selection
Individuals in one sex compete
Intersexual selection
Mate choice (usually females)
sexual dimorphism
Differences between the sexes in secondary sexual characteristics.