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Macroevolution
Broad pattern of evolution above the species level. The fossil record shows this over large time scales such as
the emergence of terrestrial vertebrates
the impact of mass extinctions
the origin of key adaptations, such as flight
A.I Oparin and J.B.S. Haldane (1920s)
Independently hypothesized that the early atmosphere was a reducing environment
Stanley Miller and Harold Urey (1953)
-Conducted lab experiments demonstrating that the abiotic synthesis of organic molecules in a reducing atmosphere was possible
-Reanalysis of molecules formed in one of these scientists’ experiments found that numerous amino acids formed under conditions simulating volcanic eruption
Abiotic Synthesis of Macromolecules
RNA monomers have been produced spontaneously from simple molecules
Small organic molecules polymerize when they are concentrated on hot sand, clay, or rock
Protocells
May have formed from fluid-filled vesicles with a membrane-like structure.
Contain properties such as replication and metabolism
Ribozymes
RNA molecules that have been found to catalyze many different reactions
EX: Can make complementary copies of short stretches of RNA
Radiometric dating
Used to determine the age of a fossil (Up to 75,000 yrs old)
A radioactive “parent” isotope decays to a “daughter” isotope at a characteristic rate
-each isotope has a known half-life (time required for half the parent isotope to decay)
Stromatolites
Rocks formed by the accumulation of sedimentary layers on bacterial mats
Date back 3.5 billion years ago
Endodymbiosis
Eukaryotes originated via this process when a prokaryotic cell engulfed a small cell that would evovled into a mitochondrion.
-An endosymbiont is a cell that lives within a host cell
-Oldest fossils of eukaryotic cells date back 1.8 billion years
The Cambrian Explosion
-Refers to the sudden appearance of fossils resembling modern animal phyla in the Cambrian period (535 to 525 million years ago)
-Provides the first evidence of predator-prey interactions
-Had a “long fuse”
Quaternary Period (0.01-2.6 mil yrs ago)
-Historical Time
-Ice ages
-Origin of genus Homo
Neogene Period (5.3-23 mil yrs ago)
-Appearance of bipedal human ancestors
-Continued radiation of mammals and angiosperms; earliest direct human ancestors
Stromatolites
-Rocks formed by the accumulation of sedimentary layers on bacterial mats
-Date back 3.5 billion years ago
Endosymbiosis
-Process in which eukaryotes originated
-Prokaryotic cell engulfed a small cell that would evolve into a mitochondrion
Theory of Plate Tectonics
Theory that Earth’s crust is composed of plates floating on Earth’s mantle
Pangaea
Formation of the supercontinent Pangaea about 250 million years ago and had many effects
Effects of Pangaea
Deepening of the ocean basins
Reduction in shallow-water habitat
Colder and drier climate inland
Organisms must adapt, move or risk extinction as the climate changes in response to continents moving toward or away from the equator
Separation of landmasses can lead to allopatric speciation
Mass Extinctions
-Can be caused by changes to a species’ biotic or abiotic environment
-Occurs when the rate of extinction has increased dramatically
The Permian Extinction
-Defines the boundary between the Paleozoic and Mesozoic eras 252 million years ago
-Occurred in less than 500,000 yrs and caused the extinction of about 96% of marine animal species
Factors contributing to The Permian Extinction
-Extreme volcanism in Siberia
-Global warming and ocean acidification resulting from the emission of large amounts of CO2 from volcanoes
-Anoxic (O2 deficient) conditions resulting from nutrient enrichment of ecosystems
Cretaceous Mass Extinction
-Occurred 66 million years ago
-Dinosaur/birds; jurassic park
-MOST RECENT
-Iridium presence in sedimentary rocks suggest METEORITE IMPACT off the coast of Yucatan Peninsula
Adaptive radiation
The rapid evolution of diversely adapted species from a common ancestor
Adaptive radiations may follow:
-Mass extinctions
-Evolution of novel characteristics
-Colonization of new regions
Reasons a Sixth Mass Extinction is Under Way:
-Many species are declining rapidly due to habitat loss, introduced species, over-harvesting, and other factors
-Climate change (higher temperatures can increase extinction rates)
Paleogene Period (66-23 mil years ago)
-Origins of many primate groups
-Angiosperm dominance increases; continued radiation of most-present day mammalian orders
-Major radiation of mammals, birds, and pollinating insects
Cretaceous Period (66-145 mil yrs ago)
-Flowering plants (angiosperms) appear and diversify;
many groups of organisms, including most dinosaurs; become extinct at end of period.
Jurassic Period (145-201 mil yrs ago)
-Gymnosperms continue as dominant plants; dinosaurs abundant and diverse
Triassic (252-201 mil yrs ago)
-Cone-bearing plants (gymnosperms) dominate landscape; dinosaurs evolve and radiate; origin of mammals
Permian (252-299 mil yrs ago)
-Radiation of reptiles; origin of most present-day groups of insects; extinction of many marine and terrestrial organisms at end of period
Carboniferous (299-359 mil yrs ago)
-Extensive forests of vascular plants form; first seed plants appear; origin of reptiles; amphibians dominant
Devonian (359-419 mil yrs ago)
-Diversification of bony fishes; first tetrapods and insects appear
Silurian (419-443 mil yrs ago)
-Diversification of early vascular plants
Ordovician (443-485 mil yrs ago)
Marine algae abundant; colonization of land by diverse fungi, plants, and animals
Cambrian (485-541 mil yrs ago)
-Sudden increase in diversity of many animal phyla (explosion)
Ediacaran (541-4600 mil yrs ago)
-Diverse algae and soft-bodied invertebrate animals appear
-Oldest fossils of eukaryotic cells appear
-Concentration of atmospheric oxygen begins to increase
-Oldest fossils of cells (prokaryotes) appear
-Oldest rocks on Earth’s surface
-Orgin of Earth