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Outer Ear
Includes the pinna, ear canal, and tympanic membrane.
Pinna
Protects the ear canal, provides sound amplification, and aids in localization.
Ear Canal
Acts as a band-pass filter (resonator system) that amplifies middle frequencies.
Tympanic Membrane (Eardrum)
Cone-shaped membrane that closes the ear canal and connects to the malleus.
Distortion
Creation of new frequencies not present in the original input.
Clipping
Occurs when sound exceeds maximum amplitude, causing the wave to be cut off.
Binaural Cues
Sound localization info from both ears (ILD and ITD).
Interaural Level Differences (ILD)
Difference in sound level/intensity between ears; prominent at high frequencies.
Interaural Time Differences (ITD)
Difference in sound arrival time between ears; prominent at low frequencies.
Monaural Cues
Sound localization info from one ear, depending on sound elevation.
HRTFs (Head-Related Transfer Functions)
The outer ear's amplitude response curve used for localization.
Middle Ear
Air-filled cavity housing the ossicular chain, located in the temporal bone.
Ossicular Chain
Connects the eardrum to the oval window (Malleus, Incus, Stapes).
Malleus
Attached to the eardrum.
Incus
The middle bone ("anvil").
Stapes
Attached to the cochlea via the oval window; the smallest bone in the body.
Middle Ear Function
To transfer energy from air to fluid, overcoming impedance mismatch.
Impedance Mismatch
Difficulty of energy transformation from air (low) to fluid (high).
Area Transformer
Main mechanism for overcoming impedance mismatch (eardrum to oval window area difference).
Stapedius Muscle
Contracts during the acoustic reflex to dampen low frequency sound.
Eustachian Tube
Connects middle ear to nasopharynx to equalize air pressure.
Otitis Media
Fluid in the middle ear; causes low frequency hearing loss due to increased stiffness.
Conductive Hearing Loss
Damage to the outer or middle ear.
Cochlea
The receptor organ for hearing; transduces mechanical energy into electrical energy.
Base (of Cochlea)
Narrower, stiffer section; processes higher frequencies.
Apex (of Cochlea)
Wider, less stiff section; processes lower frequencies.
Tonotopic Organization
Cochlea is organized by frequency (high at base, low at apex).
Traveling Wave
Wave of displacement moving along the basilar membrane from base to apex.
Scala Media (Cochlear Duct)
Central chamber filled with endolymph (+80 mV potential); contains the Organ of Corti.
Endolymph
Fluid in the scala media with a high +80 mV electrical potential.
Perilymph
Fluid in the scala vestibuli and scala tympani.
Organ of Corti
Sits on the Basilar Membrane; contains Inner and Outer Hair Cells.
Inner Hair Cells (IHCs)
Sensory transducers; send 95% of information to the brain (via Type 1 afferent neurons).
Outer Hair Cells (OHCs)
The "Cochlear Amplifier"; change length/stiffness to sharpen sound (via protein prestin).
Electromotility
The ability of OHCs to extend and contract to amplify sound.
Tip Links
Protein bridges that connect stereocilia, allowing them to move as one unit.
Stria Vascularis
The "Cochlear Battery"; maintains the endocochlear potential by supplying K+.
Depolarization
Cell becomes more positive (excitation), allowing K+ in.
Hyperpolarization
Cell becomes more negative (inhibition).
Otoacoustic Emissions (OAEs)
Sounds produced by the ear, generated by OHCs; used for newborn hearing screening.
Loss of OHCs
Leads to loss of the cochlear amplifier and distorted sound.
Loss of IHCs
Causes severe/profound hearing loss because no information can be sent to the brain.
Cochlear Synaptopathy
Disconnection between IHCs and afferent nerve fibers (e.g., from noise or aging).
Sensorineural Hearing Loss
Damage to the cochlea (inner ear) or auditory nerve.
audibility
detecting sound
intelligibility
understanding sound
tinnitus
ringing in the ears
sound waves
acoustic
vibrations
mechanical
neutral signals
bio-electric
sound
vibration of an object that causes oscillation of a medium through which energy propagates
elasticity
the ability of a mass to return to its natural shape
inertia
property to resist change
acoustical signals
variations in air pressure
mechanical signals
variations in position (displacement)
electrical signals
change in voltage
period
amount of time to complete one cycle
frequency
cycles per second
amplitude
amount of displacement
root means squared amplitude
0.707 A
phase
what is the displacement at a given time (radians)
friction
decreased motion over time; amplitude gradually decreases, frequency not impacted
pressure
force per unit area
node
where the wave hits zero
antinode
peak amplitude of the wave
condensation
region of high density and high pressure
rarefaction
region of low density and low pressure
speed of sound
350 m/s
sound pressure doubles
+ 6 dB
pressure increase by factor of 10
+ 20 dB
intensity doubles
+ 3 dB
intensity increase by factor of 10
+ 10 dB
power
rate at which energy is transformed by a wave
inverse square law
intensity is inversely related to distance
see exponential decrease as distance increases
distance doubled
6 dB decrease
absorption
sound energy is taken in by materials and converted into heat
anechoic room
designed to have no sound
reverberation time
time for reduction by 60 dB
reflection
sound bounces off a surface
interference
interaction of 2+ waves
constructive interference
two waves in phase; add to create larger sound pressure
destructive interference
two waves out of phase; add to make less sound pressure
reverberation
persistence of a sound in an enclosed space
standing wave
stable patterns of interference caused by reflections
sound shadow
area where there is little sound
complex sounds
what we hear in the real world
pure tone
single sine wave
fundamentals frequency
lowest frequency of a complex sound
octave
doubling a frequency
periodic wave
a waveform that repeats over time
aperiodic wave
a waveform that does not repeat over time; have infinite period
transience
brief pulse in music
beats
small frequency difference between f1 and f2
fourier’s theorem
any complex wave is a sine waves
fourier analysis
decomposition of a wave
fourier synthesis
reconstructing a wave
amplitude modulation
change in amplitude over time
frequency modulation
bandwidth over which the signal changes
SAM tones
2 sine waves multiplied together
gaussian noise
instantaneous amplitude probability follows a normal curve