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Biotic factor
living things w/in ecosystem
Abiotic factor
non-living components ecosystem
Abiotic factors influence on terrestrial organisms
Temperature + precipitation
Abiotic factors influence on aquatic organisms
Light + nutrients
Why are temperatures at lower latitudes higher?
Receive more solar radiation
-> Equator (0) warmer > N + S pole (90)
What causes seasons?
Earth's tilt
Why is there less precipitation at mid latitudes
- Air circulation patterns
- Descending dry air absorbs moisture
- Ascending air releases
- Warm air rises
What are climatic zones?
Regions earth defined patterns temperature + precipitation
Describe trophics
Warm, wet, weakly seasonal
Temperate zone
Highly seasonal, cold winters + hot summers
Polar
Year round low temps, 24h darkness/ light @ solstices
Curvature earth = low angle incoming sunlight
What are environmental gradients
- Steady change environmental variable through space
Latitudinal gradients
- Steady change environment across latitude
- Steeper land + N hemisphere
- Becoming more shallow cut world warming
- Temp increase as approach equator
Elevational gradients
- Oxygen availability decrease w/increase elevation as less atm pressure
- Lower pressure @ higher elevations
- Atm pressure decrease w/alt
- Limit animal dist
- Lower pressure = lower temp
Precipitation and elevations
- ppt increases higher elevations windward side mountains
1) cool air flows
2) Ppt
3) Rain shadow
- Higher ppt up windward side mountain range
- Air cools = Water vapour condenses = falls rainfall
- Descending air + reduced moisture = rain shadow leeward side = sheltered from prevailing winds
What are rain shadows?
Region little rainfall bcus shelters prevailing rain-bearing winds by mountain range
3 types Biodiversity
Genetic d in pop = diverse genome
Species d ecosystem = many species + interacting
Community + ecosystem d landscape entire region = many ecosystems
Biodiversity Scales
Alpha d = species d community/local scale (1 eco)
Beta d = species d between 2 communities/ ecosystems
-> Bio d (S1-S2) = (S1-C) + (S2-C)
Gamma d = total species d across landscape
Interactions species + ecosystem limit species distribution
Dist organisms set by eco facts + eco history = vicariance
Dispersal = organism new place
Abiotic tolerances requires species intra determine survive + reproduce
Biome
Major life zone characterised characteristic plant communities
Average conditions + pattern seasonal variation determines
Large, naturally occurring community organisms occupying major habitat
Primary productivity
Determines by temp + ppt
= Rate energy sun converted in2 biomass by photosynthetic producers
Tropical rainforest
High ppt, wet + dry seasons, temp high, highest animal + plant diversity
Desert
low ppt, temp highly variable seasonally + daily
adapt heat + desiccation tolerance
Coniferous forests
Cold winters, warm summers, cone-bearing trees, fire dependent, migratory birds + mammals
Tundra
Cold winters, cool summers, herbaceous, permafrost restricts plant growth, migratory birds
Chaparral
Seasonal, cool fall + winter + spring, hot summer temp, high diversity + endemism, small mammals
Fire + drought adapt
Elevation + Latitude + Biomes
Air temp decreases w/increasing lat
Biomes high latitudes = high elevations
Latitudinal gradient equator -> poles = tropical forest, desert, coniferous forests tundra
Niche
Combination biotic + abiotic factors species needs reproduce
Compare + contrast Chaparral and tundra biomes
- Both = low-growing herbaceous shrubs, drier
- Different ppt, climate, biodiversity
- Tundra organisms adapted cold = more fur density + fat stores, slower metabolism
- Chaparral = drought adapt = waxy cuticle, large ears release heat
Pros biodiversity
- Increased primary productivity
- Diverse pollinators = supports plant reproduction
- Soil health + nutrient cycling
- Resilience environ change
- Genetic d future
- Food security
Cultural, aesthetic, rural value
- Bio d increases NPP + resilience stress
NPP
= Net primary productivity
Total amount biomass produced by plants
Resilience
= Ability bounce back after stress
Life history
= suite traits related species' lifespan + timing + pattern reproduction
E.g. size birth, growth pattern, age + size maturity, life length, size + number sex ration, offspring, duration, investment parental care
Semelparous
Reproduces once = short adult lifespan
Interoparous
Reproduces multiple x throughout life
Survival adults high = survival juveniles
r =
rate pop increase
k =
carrying capacity = how many indies given area hold
R-selection
Selects life history maximises reproduction + ability pop increase rapidly @ low density
K- selection
Density-dependent selection
Selects LH traits enhance indies fitness when pop high (close carrying capacity) + stable
Cohort
Group Indies same age birth -> death
Life table
Age specific summary survival + reproductive rates w/in pop
Survivorship curve
Plot proportion/cohort number still alive each age + pattern survivorship pop
- Straight line = constant rate death
- log scale
Types survivorship curve
1) Low death rate early + middle life, sharp decrease later life
-> large animals, few offspring
2) Constant
-> Rodents + invertebrates
3) High dr, drdeclines survivors early period die off
-> Large number offspring b little parental care
Allocation tradeoffs
Trade off long life + diversity
Principe allocation
- Limited resources invest different activities + functions
- Growth, survival, reproduction
- Animals allocate time + energy different activities
- Plants biomass + nutrients different parts + parts carry functions
Tradeoffs
- Current reproduction + survival
- Current + future reproduction
- Number + size offspring
- Somatic maintenance = energy spent maintain body
Explain why mice often abandon their young
- carrying young substantial time, energy, resources
- food shortages = less resources allocated
- less resources available survival + reproduction = trade off
- Survival + future reproduction over current
- Interparous = invest more somantic maintenance
Reasons modelling population growth
Conservation, resource needs, policy, max harvests, biological invasions, disease spread
Calculating population size
Nt+1 = Nt + I - D - E
Nt = number indies pop @ 1x
Calculating population size of closed population
Nt = Nt + B - D
Assume no immigration/emigration
Rate population growth
^N/ ^t = B - D = R
Exponential pop growth
- Geometric growth
- intrinsic rate increase (r) = % change pop size per capita
- r constant = pop grow exponentially
- Assumes time intervals infinitely small, continuous demography
- r>0 = pop size grow
- r = 0 same
- r < 0 = decline
- looks linear log-scale
Causes exponential growth
- Pop introduced new environment
- Catastrophic disturbance
- Important predator removed
Density-dependent population growth
- Changed br + dr w/density pop
- Exponential growth = brings + dr assumed constant regardless pop density
- Denser pop = decrease br + increase dr
- Less resources indivs
- More competition, disease, predation risk
-Logistic (S-shaped) pop growth curve
S-shaped growth curves
Density-dependent pop
- Pop low density = growth exponential
- Pop size increase = d-d factors decrease birth/decrease death
- Pop growth slows until carrying capacity
Density Independent pop growth
= Rate growth pop @ any instant limited something unrelated size pop
- External environ aspects = cold winters, droughts, storms, volcanic eruptions
- Erratic pop growth patterns
Logistic Growth Equation
dN/dt = rN (k-N/K)
rn= exponential growth model
() = linear decrease towards carrying capacity
r = intrinsic rate increase
k = carrying capacity
Competition
Occurs 2 indivs share resource + consumption 1 reduces availability others = reduced growth, survival,/ fecundity
Intraspecific competition
between indivs same species
Mechanism behind density-dependent pop growth
Interspecific competition
between indivs different species
Food, mates, pollinator, space, nests water
Exploitative competition
= comp mediated by consumption/shared resource
- Indivs/species not physically encounter
- indirect use shared resource
- 1 competitor consume = reduce supply
Inference competition
involves direct, physical interaction
1 competitor expend energy inhibit access resource other
Scarcity
no resource unlimited quantity = competition
Parasites
Derive nourishment from host = damaged/killed
Parasitoids
Lay eggs + larvae consume prey
Can herbivores be considered predators
Eat seeds + embryonic plant fully eaten
Mutualistic if seeds not digested + dispersed
Ectoparasites
live outside body
endoparasites
live inside body
Mutualism
= Both interacting species benefit increased fitness
- Direct/indirect
- conservation = seed dispersal, mycorrhizae, pollination
- increase reproductive revival, rate pop growth/size
Commensalism
1 species benefits, other unaffected
E.g. birds eat insects off buffalo
Symbiosis
Organism living in/on another organism regardless outcome
Ecological effects species interactions
Ecological processes = determine abundance, range/distribution, timing + activity interacting partners
(+) increase abundance. extend range, select synchronised activity partners
(-) oppo, select altered timings
Cycle ecological change
Ecological change -> Alters selective pressures pop -> evo change -> alters outcome eco interactions
Co-evolution
= Reciprocal evolution 2 interacting species
- Change traits indivs 1 pop response trait indivs other pop
- From competition, exploitative mutualistic intrxs
Population cyclic
- Pred overexploit prey = cyclic fluctuations
- Lag prep pop decline + red response
- Fred alters selective pressure prey = evo change
Consequences cyclic populations
- Competition decreases fitness both interacting species = competitive exclusion weaker species
- Evo act reduce competition = niche partitioning + character displacement
Competitive exclusion
= Species compete 1/+ species extinct
- Species exhibits logistic (s-shaped) growth flask w/other species
- Failure co-exist = competition same resources
Gause's principle
= 2 species competing same limited resource, species uses resource more efficiently eventually eliminate other locally
- slight advantages built into species over time
- only valid if single resource + vary time/space
Selective sweep
= Allele fixed population
Niche
= Range abiotic + biotic factors species needs survive
When does competitive exclusion occur?
Competition asymmetric + niches completely overlap
Sympatric
= species/populations overlapping ranges
Allopatric
= species/populations separated by barrier
Character displacement
= tendency characteristics (physiology, behaviour, morphology) diverge more sympatric (geographically overlapping) compared allopatric populations reduce competition
- decreases competition
Fundamental niche
Resources used/conditions tolerated absence competitors
Realised niche
Resources used/conditions tolerated when competition occurs
Niche overlap not complete
Weaker competitor uses non-overlapping resource
Niche differentiation/resource partitioning
= Evolutionary change use by competition over generations
- decrease niche overlap + amount comp = coexistence more likely
Outcomes exploitative interactions
- Boom + bust pop cycles
- decrease abundance + range prey/host
- Defensive + offensive adaptations morph, physiology, behaviour
Defensive adaptations
- Eco selection pressure = adaptation
- Evo response = m/p/b adaptations = prevent detection + eaten
- Plants = mechanical + chemical
- Host = immune system, sequester toxins, behavioural
Offensive adaptations
m/p/b = eat + find prey
Parasite adaptations
- Disrupt immunity
- Manipulate host chem
- Physical structures stay in/on host
- Manipulate host behaviour = facilitate transmission + increase survival
-> ants climb blades grass = eaten sheep = next host
Outcome mutualistic interactions
Increase abundance + range interacting partners
Adaptations promote interactions
Outcome species interactions
(-) Decrease br + increase dr
(+) opposite
- Impacts pop demography + species range limits
- Natural selection enhances traits = decrease -ve itrxs = co-evolution interacting partners
What limits primary productivity?
Water, energy, nutrients (N, P, iron, salt)
What drives productivity gradients?
solar radiation
Main stages water cycle
Evaporation, precipitation, runoff
Stocks
Stores/pools matter
10^3km^3
Fluxes
Movements between pools
km^3/yr
Nitrogen containing molecules
DNA, ATP, RNA, nucleic acids, proteins
Chlorophyll, rubisco