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hominoid
superfamily: hominoidea
includes: lesser and great apes
hominid
family: hominidea
includes: humans, their fossils ancestors, Pan (Chimpanzee), Pongo (Orangutan) and Gorilla
hominin
tribe: hominini
includes: humans and their fossil ancestors
biases in the fossil record
sampling errors
differential preservation - only a handful of organisms that die are in the right condition to be fossilized
taphonomic conditions (size and makeup of animals)
gaps in the fossil record
anagenesis
gradualism (straight line evolution)
lineage gradually changes over time
cladogenesis
punctuated equilibrium
split into two distinct species
ex: new areas/environmental change
gradualism
small changes over generations
transitional forms (mix of ancestral and primate characteristics)
punctuated equilibrium
long periods of stasis, short periods of rapid change
no transitional forms
relative dating
comparatively younger and older
absolute dating
exact age in years
stratigraphy
relative dating method
analyzing the sequence of rock layers (strata) to determine relative ages.
law of superposition
lower (bottom) stratum older than higher (top) stratum
fluorine analysis
relative dating method
fluorine builds up, exposure to groundwater
more fluorine = older fossil
occurs slowly
only used on fossils from the same area
can’t compare different sites
faunal correlation/biostratigraphy
relative dating method
trata from multiple locations compared
index fossils - fossils that have a wide geographical distribution but existed for a short period of time (identifies stratigraphy/time lines) → both absolute and relative dating methods
thermoluminescene
absolute dating method
electron release associated with heating crystalline rock
dendrochronology
tree ring dating
radioactive decay: carbon-14 (radio carbon)
absolute dating method
method used to determine the age of organic materials (once-living things) by measuring the amount of a radioactive isotope of carbon, carbon-14 (14C), remaining in the sample
determines half life (amount of time it takes for half of the unstable isotope to decay into the stable isotope): 5730 years
reliable ~ 50 KYA
radioactive decay - argon
absolute dating method
determines the age of geological materials, particularly volcanic rocksK/Ar dating (^40K decays into ^40Ar or ^40Ar decays into ^39Ar)
Half life: 1.3 MY
Fossils: 100+ KYA
cenozoic era
65 MYA - present
tertiary period
65 MYA - present
paleocene
65-53 MYA
eocene
53-25 MYA
oligocene
35-23 MYA
miocene
25-5 MYA
pliocene
5- 1.8 MYA
pleistocene
1.8 MYA - 10 KYA
holocene
10 KYA - present
arboreal hypothesis
primates emerged due to adaptations to life in trees
visual predation hypothesis
primates emerged due to hunting in trees
angiosperm radiation hypothesis
primates emerged due to fruit eating n trees
paleozoic era significance
first vertebrates: 500 MYA
mesozoic era significance
placental mammals: 70 MYA
cenozoic era significance
mammals diverge: 65 MYA
era with first appearance of primates
paleocene (65 MYA)
primates associated with paleocene
infraorder plesiadapiforms (Carpolestes, Purgatoriidae, Palaechthonidae, Paromomyidae, Picrodontidae, Plesiadapidae)
infraorder plesiadpiforms
proprimates (early primate ancestors)
similar to modern primates
primitive traits: no postorbital bar/convergent eyes, lacked opposability and retained claws, small brain and specialized rodent like teeth)
~ 60 MYA: Western North America, Europe
carpolestes
wyoming (58 MYA): tropical forest
primate features: grasping feet, nails
proprimate features: claws, nonconvergent eyes
eocene primates
euprimates (earliest true primates)
adapids, omomyids, basal anthropoids
primitive traits: nails, grasping hands and feet, opposable thumbs and toes, reduced olfaction, increased vision (postorbital bars), larger brains
omomyids
tarsiers
traits: short face, V-shaped lower jaw, large eye orbits, long legs
nocturnal
vertical clinger and leaper
adapids
lemurs and lorises
traits: dental comb, long legs and tails, grasping feet
diurnal
vertical clinger and leaper
basal anthropoids
eosimias (china 45-40 MYA): short heel, anthropoid like teeth
biretia (egypt 37 MYA): teeth anthropoid like
oligocene primates
- early anthropoids
parapithecids (adium)
oligopoithecids (basal anthropoids)
early catarhhines
aegytopithecus
saadinus
primate traits: post orbital plate, expanded brain, 2.1.2.3, sexual dimorphism
generalized arboreal quadrupeds
larger body sizes
aegytopithecus
ancestral to old world monkeys and apes
arboreal quadrupeds
early catarrhine
35- 55 MYA
saadinus
precedes old world monkeys and ape split
early catarrhine
29 - 28 MYA
mioecene primates
proconsulids, dryopithecids, sivapithecids, gigantopithecus, african apes (ouranopithecus, nakalipithecus, choropatihecus, simiolus mintus)
apes
HOMINOIDS!
traits: large complex brains, generalized skeletons, generalized dentition (y-5 molars, frugivores/folivores)
first old world monkey
kenya (15 MYA)
pronconsulids
22-17 MYA
primitive traits: v shaped lower jaw, longer legs
hominoid traits: y-5 molars, sectorial premolar, frugivore molar, no tail
dryopithecids
18-15 MYA
europe
expansion OUT of africa
proconsular teeth
modern ape bodies (suspensory behavior)
apes in asia
sivapithecids
gigantopithecus
sivapithecids
12-8 MYA
arboreal quadruped?
orangutan like skull
proconsulid like body
gigantopithecus
8-5 MYA
largest known
african apes
occurred in late miocene
cooling and drying trend → grasslands and woodlands appear
ouranopithecus
nakipithecus
chorapithecus
last common ancestor? (11-10 MYA)
similous mintus
small bodied ape
folivore
12.5 MYA
pliocene - description
5 - 1.8 MYA
earliest definite hominins
diversification of old world monkeys
dramatic loss of hominoid diversity
pleistocene - description
1.8-10 KYA
modern non human primates
modern humans and other hominins
maybe primates, maybe not
paleocene
age of prosimians
eocene
age of monkeys
oligocene
age of apes
miocene
age of hominins
plio-pleistocene
modern day
holocene
human’s last common ancestor epoch
mioecene
plesiadapiforms
archaic primates or primate like placental mammals
anthropoids
monkeys, apes, humans
plesimorphic
having features shared by different groups with a common ancestor/ancestral traits
Hominins vs Apes
obligate bipeds
reduced canines
flat faces
larger brains
earliest hominins
sahelanthropus tchadensis, orrorin tugenesis, ardipithecus ramidus, keyanthropus platyops
sahelanthropus tchadensis
chad, africa 7-6 MYA
350 cc, foramen magnum, prominent brow ridges, small canines
earliest hominiin
“toumai” fossil
orrorin tugenesis
tugen hills, kenya: 6 MYA
thick enamel, large canines, femur at a neck angle
ardipithecus ramidus
ethiopia: 4.4 MYA
numerous fossils
fairly complete “ardi” skeleton
opposable hallux, upper limbs, thin molar enamel, small canines, anterior foramen magnum
kenyanthropus platyops
kenya: 3.5 MYA
small canines, thick enamel, flat face
contemporaneous with australopithecus
hominin timeframe
mioecne - pliocene (6-2.6 mya)
robust
paranthropus/robust australopith
gracile
australopithecus
robust/gracile divisions represent
chewing adaptations
paranthropus and australopithecus
4-1 MYA
primitive traits: ape like skull and body size, small cranial capacity
bipedal
paranthropus
paranthropus aethiopicus, boisei, robustus
east and south africa
2.8-1 MYA
no evidence of tools
high degree of sexual dimorphism
coexisted with gracile forms and early Homo
paranthropus traits
4-5 feet tall
450-550 cc
omnivores with fall back foods
megadontia (large molars/premolars), thick molar enamel, non projecting canines
pananthropus aethiopicus
black skull
west lake turkana, kenya
2.5 MYA
410 cc
ancestral to other paranthropus
paranthropus boisei
“nutcracker man”/”zinj
olduvai gorge,tanzania
1.75 mya
500-559 cc
larger cheek teeth, flatter skull, larger brain, thickest enamel
first to use stone tools?
paranthropus robustus
kromdraai, south africa
2-1.2 mya
large teeth, adapted to dry enviroment, sexual dimorphism
australopithecus
afarensis, africanus, garhi, sediba
evolved into homo, coexisted with robust forms and early homo
east and south africa
4.2-1.8 MYA
mostly vegetarian
high degree sexual dimorphism
400-500 cc (smaller brains)
small bodies
moderately large molars, intermediate dental arcade
bipedal and arboreal
australopithecus africanus
south africa
400-500 cc
small dull canines
farmen magnum
taung child (2.8 MYA)
mrs. ples (2.5-2.1 MYA)
australopithecus afrarenis
ethiopia
small canines, bipedal, curved fingers
430 cc
lucy: 3.2 MYA (1 meter tall)
Dikika Child/Lucy’s Child/Selam: complete skeleton
laetoli footprints
tanzania
3.6 MYA
volcanic sediments show bipedal locomotion of Australopithecus Afrarensis (3 individuals)
Australopithecus tool use
fossilization
dikika: cut marks (naturally sharp rocks, scavenged for meat)
paranthropus boisei found near stone tools
australopithecus garhi
2.5 MYA
link betweem australopithecus afrarensis and homo
awash and ethiopia
australopithecus sediba
malapa cave, south africa
~ 2 MYA
420-450 cc
bipedal
mosiac features (transitional species between australopithecus africanus and homo)
piltdown man
1912, evolutionary link between apes and humans
FRAUD
bipedalism
maintaining balance on the stance leg while in swing
habitual bipedalism
bipedalism as a primary and most efficient form of locomotion (hominins)
obligate bipedalism
committed to bipedalism, cannot locomote efficiently any other way (humans)
anatomical adaptations to bipedalism
changes in foramen magnum position
changes in vertebral column, thorax, hip, femur tibia, and foot
foramen magnum - humans
anterior posture (front)
foramen magnum - non human apes
posterior position (back)
back and thorax - human
s-shaped spine, barrel shaped chest
back and thorax - non human apes
c-shaped spine, funnel shaped chest
pelvic shape - humans
shorter and broader, bowl shaped, stabilizes weight distribution
pelvic shape - non human ape
long narrow pelvis, positioned along lower spine
abductor muscles - humans
gluteus medium and minimus (stabilizes leg)