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When did the first angiosperms evolve
130 MYA
when did grasses evolve
30-50 MYA
what delayed the movement of plants onto land
avoiding desiccation - not drying out
when did first land plants evolve
500 mya
primary growth
occurs in meristematic tissues
secondary growth
uses two lateral meristems, the vascular cambium and cork cambium
roots
anchorage, uptake of water and mineral ions
stems
support, nodes for growth points, vascular tissue
leaves
gas exchange, vascular tissue
cell wall
provides immense strength to plants
chloroplast
each cell has can have several, where photosynthesis takes place
large central vacuole
used for storage and ion/water absorption; plays major role in cell turgidity
plastids
unique organelles in plants
plasmodesmata
produces a link between cytosol of neighboring cells that allows for solute transfer
vacuole
occupies 50-98% of the cell
tonoplast
membrane that surrounds the central vacuole
storage compounds in vacuole
ions, sucrose, and secondary metabolites that maintain cell turgidity
primary cell wall
thin and flexible layer that is mostly cellulose
secondary cell wall
rigid and provides support, more lignin
middle lamella
the glue that binds cells together
gametophyte
generation that begins with meiosis
sporophyte
generation that begins with fertilization
gravity
weak force but governs the entire universe
electromagnetic
the attractive or repulsive force between electric charges and magnets
strong nuclear
exist to hold protons and neutrons together
weak nuclear
can change a neutron to a proton -> changes element
electromag, w. nuclear, and s. nuclear
all connected together
1st law of thermodynamics
Energy cannot be created or destroyed
2nd law of thermodynamics
measure of the capacity of a system to undergo change, the entropy of the universe is always increasing, and the universe trends towards chaos and disorder
3rd law of thermodynamics
the law that states that in order for motion to stop we have to reach absolute zero
Gibbs free energy
biological rxns require energy and total energy = G
Gibbs free energy equation
ΔG = ΔH - TΔS
exergonic rxn
energy releasing; K > 1
endergonic rxn
Energy requiring; K < 1
spontaneous rxn
G is -
nonspontaneous rxn
G is +
oxidation
addition of O, removal of H, removal of e, displacement of e
reduction
removal of O, addition of H, addition of e, displacement of e
photosynthesis
a series of Redox rxns (water oxidized, CO2 reduced)
enzymes
modified proteins that aren't changed by rxn
enzyme catalyzed rxns
high specificity, high rxn rates, mild rxn conditions, opportunity for regulation
unique properties of water
universal solvent, polarity, high specific and latent heat, high cohesion, high adhesion, high surface tension
universal solvent
results from size and polarity
polarity
H-O-H covalently bonded, opposite bonds resulting in H bonding
high cohesion
polarity results in attraction between H2O; high tensile strength permits capillary action
high adhesion
attraction of water to solid phase
high surface tension
water prefers water not other gasses, energy is required to incr. surface area of gas-water interface
diffusion
Movement of molecules from an area of higher concentration to an area of lower concentration over a short distance (cellular)
mass flow
movement of groups of molecules in response to a pressure gradient (environmental gradient)
osmosis
Diffusion of water through a selectively permeable membrane
water potential
free energy of water per unit volume (Jm-3 or Pa)
water potential equation
Ψw = Ψp+Ψs+Ψg
osmotic potential Ψs
more solutes = less energy; dilution effect; only ever negative, Ψs = 0 then pure water
hydrostatic pressure potential Ψw
can be + or -, + = turgor, - = tension
gravitational potential Ψg
grav. is true force acting on water, increase height incr. Ψg by 0.1 MPa per 10 m
turgor
pushing potential (+)
tension
pulling potential (-)
turgor potential
has greater potential decline w/ initial decr. in cell volume
elastic modulus
the slope of Ψp - V curve in cells
less water/turgidity
higher ion conc and lower Ψs
pressure potential
directly linked to osmotic potential
dessication
some species can recover, but most can't due to tears in membrane and inability to recover cell contents
plasma membrane
semipermeable membrane that regulates molecule and ion movement in and out of cell
phosopholipid
hydrophilic head and hydrophobic tail
bilayer structure
results in a highly fluid, very stable, and impermeable membrane
transport
movement of molecules and ions across a membrane or w/in or between cells
passive transport
spontaneous movement down a potential energy gradient (diffusion/osmosis/mass flow) and proceeds until equilibrium
active transport
against conc. gradient, not spontaneous and requires energy
ATP synthase
enzyme that uses protons to perform ATP synthesis
proteins
membranes can have >= 50%, have lateral fluidity
primary role of membrane proteins
metabolism, transport, energy transduction
transport proteins
channels, carriers, and pumps
channels
passive transport, can open and close. Low pH—> high pH
carriers
switch between active and passive transport (anti- and sym- porters). PMF
pumps
active transport that move ions against gradient. High pH—> low pH. Atp hydrolysis
confirmational binding
change of structure of carriers and pumps depending on the molecule theyre transporting
proton motive force
secondary active transport, conc. of H outside the cell creates an electrochemical energy and pH gradient
symporters
both ions/molecules travel in the same direction (PMF)
antiporters
ions/molecules move in opposite directions (PMF)
vacuole
takes up ions so they don't interfere w/ organelles; water follows ions to help maintain cell turgidity
cytosol
has a neutral pH to ensure proper organelle function; have higher conc. of sugars, amino acids, and metals to be available for use
aquaporins
channel proteins that facilitate water movement across membranes (gated channel), allows water to move much faster than just diffusing across membrane
slowest -> fastest membrane transport
pump -> carrier -> channel
apoplast
cell wall space between adjoining cells (5-20% of plant)
symplast
cell-cell transport between plasmodesmata
Casparian strip
a barrier to all apoplastic movement and water/ions must be allowed into symplast to continue forward. keeps water from experience reverse flow in drought conditions.