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The Goal of the Sensory System
Physical Stimuli and Psychological Experiences
Physical Stimuli
Wavelengths of light, Sound Waves, Chemicals, and Pressure
Psychological Experiences
Smelling, seeing a painting, or holding someones hand
The Three Somatosensory Systems
Nocioception
Hapsis
Proprioception
Receptors
Refers to individual neurons that control the physical world around us into action potentials
Nocioception
Pain, temperature, itch
Hapsis
Fine touch and pressure
Proprioception
Body Awareness
How does pressure on our skin get translated to a neural signal?
The displacement of skin causes the opening of sodium channels, which leads to depolarization and then ACTION POTENTIAL
How is speed determined?
By myelin and diameter
How does diameter determine speed?
The larger the diameter of the axon, the faster the rate of action potential (A-beta is faster than C fibers that have no myelin and a small diameter)
How does myelin determine speed?
it speeds up the rate in which the AP is sent all the way down the length of the axon
How does information from the surface of the skin get to the spinal chord?
1. Starts at the skin surface which causes an AP in the receptor
2. AP is propagated all the way down to the axon
3. Travels through dorsal pathway and into the spinal chord
Bipolar Neuron
You can see the cell body, and resides in a collection of cell bodies called the DORSAL ROOT GANGLION
The Hapsis System
Responsible for touch and pressure
Topography
Orderly entry of information from dermatomes into the spinal chord
Example of Topography
When touch happens at the foot, enters the spinal chord at very low levels.
Touch to shoulder, enters at a much higher level.
Dermatomes
Large regions of skin. Who's neurons travel in the same peripheral nerve, with cell bodies in the same dorsal root ganglion and enter the spinal chord at the same level
The Thalamus
Is carved into small subregions for specialization in particular types of sensory and motor functions
From the Skin to the Brain
1. Information enters via the Dorsal-root ganglion
2. Information projects to and synapses on to cells of the dorsal column nuclei
3. Contralateral projection to ventral posterior nucleus of the. thalamus
4. primary somatosensory
How does Somatosensory get to the Thalamus?
It travels through the ventral posterior nucleus of the thalamus
Where is the primary somatosensory cortex?
Parietal Lobe
Responsibility of the somatosensory cortex?
Responsible for perceiving touch to the surface of the skin
Central Sulcus
separates frontal and parietal lobes, posterior to it is the PSC
What is the homunculus?
It is a somatosensory map
How does a somatosensory map work?
Left side process right side of body
Right side process left side of body
What are receptive Fields
Region of physical space where it is possible for a physical stimulus to alter the firing rate of the neuron being measured
Center of Receptive Field
increases firing rate
Touching the surround of the receptive field
decreases firing rate
Touching Center and Surround
cancels the responses out
touching the outside of the field
no change - not within its field
Plasticity
Brain Structures can change to better cope with the environment
Motor Cortex Axons
Axons from this do not innervate muscles directly
What do motor cortex axons do if not innervate muscles directly?
Their either directly or indirectly innervate neurons of the ventral horns of the spinal chord
Spinal Motor Neurons
Neural elements that ultimately talk to muscles and gets them to contract
Motor Unit
Single spinal motor neuron and all of the individual muscles fibers it contracts
Muscle Spindle Neurons
Status of a muscle is monitored by receptor systems
When do muscle spindle neurons increase firing rate?
when the muscle is stretched
Where are muscle spindle neurons located?
They are embeded in the muscle itself and report the length of the muscles
Wavelength determines....
Color
Amplitude determines...
Brightness
Sympathetic Activation and the Iris
Sympa. is the activation which leads to pupil dilation
Parasympathetic Activiation at the Iris
Activation of the Parasympa. leads to pupil constriction
Sensitivity (Pupil)
The larger the pupil size, the more light permitted in.
Widening the pupil increases sensitivity
Acuity (Pupil)
In bright light, the pupil constricts.
It improves acuity
Binocular Disparity
Images on the retina are 2D BUT with the image on each retina offset (BD) we construct a 3D psychological image
The Retina
is a 5-layered sense organ for vision
What is on the Back layer of the Retina
Receptors for light (rods and cones)
What is on the front layer of the Retina
Retinal Ganglion Cells
Axons travel out of the eye and to the brain
Two problems with the Retina due to neuron arrangement
1. Light is filtered and distorted on its way through the neuronal cell layers to the receptors
2. Axons of the retinal ganglion cells ultimately have to cross through the retina to make it out of the eye
Solution to light distortion
Fovea
What is the fovea and what does it help solve?
Is a specialized area of the retina with a thinning of the retinal ganglion cell layer.
High acuity (seeing things clearest when looking directly at them), this is due to the image falling on the fovea.
Solution to axons passing the retina?
completion
What is completion?
Where the visual system uses information from the receptors around the blind spot to compete broken images
Receptors for the eye
Rods and Cones
What is a Rod?
Rod-mediated vision dominates in dim light
more sensitive to light than cones
poor acuity
lack of color
What are cones?
Cone-mediated vision dominates in ample light
high acuity
color
How to stop rods from firing action potentials?
the bleaching of rhodopsin
What does stopping action potentials do?
Stopping action potentials, or the inhibition, starts a string of effects on downstream neurons that lead to excitation of retinal ganglion cells
What do retinal ganglion cells do?
They carry information out of the eye and to the brain
How do axons from the retinal ganglion cell travel?
They travel via the optic nerve
Crossing visual information
Information from the left visual world is processed by our right visual cortex and vice versa
Where can we think our retina is divided?
At the fovea!
What are the two hemiretinas?
Nasal and temporal
What does the nasal hemiretina see?
from one eye, it sees a different part of the visual world than the temporal hemiretina from the same eye
Where do nasal hemiretinas cross?
at the midline
Do temporal hemiretinas cross at the midlinee?
NO
Where and how is information from the retina projected?
It projects to the lateral geniculate nucleus which then projects to the primary visual cortex
Retinotopic map maintains...
Order of the parts of the image are maintained due to this
Where is the Retinotopic Map located?
the lateral geniculate nucleus and primary visual cortex
Visual receptive field
an area of visual space where it is possible for light to change the firing rate of a neuron
Doral Stream is where....
information flows to the posterior parietal cortex
Ventral stream is where...
information flows to the inferotemporal cortex
What is the dorsal stream important for?
for determing where the stimuli are in space
What is the ventral stream important for?
for identifying what objects are
What is the physical stimulus for audition?
Vibrations of air molecules (sine waves)
Vibrations into neural signals
sound vibrates the tympanic membrane which vibrates the ossicles which then causes the stapes or stirrup to vibrate the oval window of the cochlea
Organ of Corti
Sense organ for audition
Chock full of hairs
Organ of Corti, the receptors for sound are hair cells
how do ion channels open up in Corti?
the deflection of hair cells in one direction causes ion channels to open
Depolarization of a hair cell causes what?
neurotransmitters to be released from it onto the next neuron and causes Action Potentials to be generated
What causes corti to be stimulated?
tones of different frequencies
Tonotopic Map
regions that respond best to one particular tone are neighbored by regions that respond best to a tone of a slightly different frequency
Where is the tonotopic map preserved to?
the cortex
Which region of the thalamus processes auditory information?
the medial geniculate nucleus of the thalamus
Primary auditory cortex
located within the temporal lobe of the cortex
it is organized in columns
Sense organ of the vestibular system?
Semi-circular Canals (SCS)
Where is the Semi-circular canal (SCS) located?
Located next to the cochlea
What is the SCS important for?
Important for balance
Translates head movement into neural signals by the movement of fluid in the SCS and stimulation of hair cells
Where does information from the hair cells of SCS travel?
travels along the cranial never VIII - the auditory nerve
How is SCS information processssed?
By the brainstem as well as the cerebellum
Which system does the SCS work closely with?
works with the visual system
Top-Down Control
Higher order areas of the nervous system (e.g. the cortext) alter sensorys and motor processing in the peripheral nervous sytems