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4.4 billion years ago
when did life begin
Endosymbiosis
symbiosis in which one of the symbiotic organisms lives inside the other.
Georges Cuvier
fossils and extinction
catastrophism (catastrophe leads to evolution and new species)
Wrong that function determines form
Jean-Baptiste Lamarck
Lamarckian evolution (Wrong): aquired characteristics are inherited and passed down. inheritance is non random (wrong)
Alfred Russel Wallace
supported Darwin's claims by determining that species were separated by natural barriers. (line between Asia and Australia.
charles darwin
toured the Galapagos Islands and developed the basis of evolution. Survival of the fittest and descent with modification.
Importance of Galapagos Islands
low biodiversity, very isolated, and unique organisms. (simplified)
Genetic forces of Evolution
Bottleneck, founder effect, non-random breeding, mutation, and variation
evidence of evolution
similar structures (5 fingers and tetrapod limb) chordates similar embryonic structures.
Embryonic Chordate Characteristics
Pharyngeal pouches, dorsal hollow nerve cord, notochord, postanal tail, and an endostyle/thyroid gland
paedomorphism
The retention in the adult of features that were juvenile in its ancestors
analagous characters
Physical structures with similar function but different underlying structures. (derived independently)
symplesiomorphy
traits unchanged from an ancestor that are inherited by a group of lineages (backbone in chordates)
autapomorphy
traits that are different than ancestors and unique for a particular lineage (helps determine differences in lineages but not link evolutionarily)
synaptomorphy
traits that are different from an ancestor and shared by more than two lineages (forelimbs nor present in sharks or fish but in present in whales, humans, cats, lizards etc.)
convergent evolution
Process by which unrelated organisms independently evolve similarities when adapting to similar environments
divergent evolution
when two or more species sharing a common ancestor become more different over time (to fill different niches)
species
individuals of a population who look similar, occupy the same niche, have the same genetic history, and the potential to interbreed and produce viable offspring in a place and time period.
speciation mechanisms (prezygotic)
habitat isolation
temporal isolation (timing)
behavioral isolation
mechanical isolation (size and shape)
genetic isolation (sperm egg protections)
speciation mechanisms (postzygotic)
Genes very different = no offspring
genes very similar = offspring (viable or not viable) and inbreeding (maybe offspring maybe viable)
vagus nerve
the tenth cranial nerve that innervates digestive organs, heart and other areas
homologous characteristics
similar in two or more species because they are inherited from a common ancestor
Carolus Linnaeus
developed binomial nomenclature, father of taxonomy
phylogenic classification
classification of organisms based on their assumed evolutionary histories and relationships.
Systematics
study of the diversity of life and the evolutionary relationships between organisms
cladograms
a branching diagram showing the cladistic relationship between a number of species.
Clade
A group of species that includes an ancestral species and all its descendants.
Apomorphy or derived character
different from the ancestral condition; evolutionary novelties
synapomorphy or shared derived character
shared by two or more species or taxa and has originated in their most recent common ancestor
Plesiomorphies
ancestral states; not informative in constructing phylogenies
Symplesiomorphies
shared ancestral states (not informative about phylogenetic relationships)
Heterotrophic
Organisms that obtain their nutrients or food from consuming other organisms.
Chordate
animal that has for at least one stage of its life: a dorsal, hollow nerve cord, a notochord, a segmented muscular postanal tail; Endostyle, and pharyngeal pouches
Subphylum Vertebrata characteristics
development of neural crest/skull
proper brain (3 parts)
Hox genes (segmented body)
vertebral column
almost all notochord develop into disks
myxini (hagfishes)
Produce slime for defense
Jawless (form a knot to feed)
Eats sporadically (snake like)
100 to 1 female to male ratio
No larval stage (unlike lampreys)
Petromyzontidae
eel-like
jawless
marine and freshwater (sea lamprey is anadromous (migrate upriver to spawn)
larval and adult form
only half parasitic
Petromyzontidae larvae stage
Ammocoetes (larvae) are benthic, filter feeders up to 7 years
petromyzontidae adult stage
Non parasitic adults (dont feed, only mate, up to 6 months)
parasitic adults (1-3 years)
Conodonta
(extinct)
soft eel-like bodies
jawless filter feeders
known for conodont elements (very small teeth)
Pteraspidomorphi
(extinct)
armored jawless fishes
extinct in late Devonian
Ostracoderms
earliest agnathans, armored, very abundant in their time period, jawless filter feeders,
Agnathans
jawless vertebrates
Gnathostomes
jawed vertebrates
Jaws
what evolutionary step is considered most important for vertebrates?
Cambrian explosion
rapid diversification of species at the beginning of the Paleozoic with the advent of jaws
placodermi
(extinct)
first jawed fishes
armored
teeth (bony plates that were not replacable)
Chondrichthyes (physical characteristics)
cartilaginous skeleton (no ribs)
paired eyes, pectoral, and pelvic appendages
most heterocercal tail (downward force)
lipid filled liver (maintains buoyancy)
placoid scales (reduce resistance)
constant teeth replacement (in lines)
Chondrichthyes (senses)
Good vision (color) (tapetum Lucidum)
good hearing,
great smell (can detect proteins, amino acids, and blood)
Electroreceptors (ampullae of Larenzini)
Lateral line (move detection)
tapetum lucidum
eyes shine, reflects light into the retina a second time)
Ampulae of Lorenzini
-sensory vesicles that respond to changes in temperature, water pressure, weak electrical fields, and salinity
lateral line
sensitive receptor system that enables fish to detect gentle currents and vibrations in the water
saltwater bony fishes (osmoregulation)
actively excrete salts, tend to gain salt lose water, always drinking
hyposmolal (hypotonic) body fluid less concentrated salt than surrounding)
freshwater bony fishes (osmoregulation)
active transport of salt in (though gills), tend to lose salt and gain water, lots of urine to expel water
Hypresomal (hypertonic) higher salt concentration than surrounding.
Hagfish osmoregulation
Iso-osmotic to their saltwater environment. No net influx or efflux of water. Similar mineral concentration to the environment. Perform as osmoconformers instead of osmoregulators.
chondrichthyes osmoregulation
keep udea in body to balance osmotic pressure
urea toxic so must produce TMAO to protect proteins.
Stenohaline
organism does not move among water salinity concentrations
euryhaline
organism can reverse water concentrations as they move between them
Halocephali
chimeras and ratfishes
deepwater, fairly unknown
venomous dorsal spine
Elasmobranchii superorders
Galeomorphi:
Squalimorphy:
Batoidea:
elasmobranchii
sharks skates and rays
diverse reproduction
age at maturation 6-18 years
hyostylic jaw suspension
most ectothermic
can be parthenogenic
parthenogenesis
Asexual reproduction in which females produce offspring from unfertilized eggs.
Hyostylic Jaw Suspension
jaw not attached directly to cranium and can be extended out
elasmobranchii reproduction
Oviparity (egg laying - 40%)
Live birth (60%)
uses yolk (ovoviviparity)
ovoviviparity and eats sibling (embryophagy)
ovoviviparity and eats ovulated eggs (oophagy)
ovoviviparity and then yolk sac attached to placenta (uterine viviparity)
batoidea
skates and rays
dorsoventrally compressed
eyes and spiracle dorsal
euhyostyly (jaw nor conected to cranium
skates
2 pelvic fin lobes, egg laying (oviporus), no stinging spike, prominent dorsal spine
squalimorphi
shark like
no anal fin
primitive most deep sea sharks
Lanternshark, cookiecutter, greenland, and megamouth sharks
galeomorphi
Shark sharks
top predators (some filter feeders)
heterocercal caudal fin
anal fin
5 fils
galeomorphi examples
great white, megalodon (extinct), whale shark (largest), Basking shark (small brain, second largest), frilled shark, great hammerhead, bull, tiger.
big three
bull shark: can enter fresh water
great white; largest
tiger shark; most aggressive
Nuctitating membrane
chandrichthyes dont have eyelids but mebranes that cover eyes from inside corner)
buccal pumping
A form of breathing in which animals take in water or air into their mouths, then raise the floor of the mouth, creating a positive pressure that pumps water or air across the gills or into the lungs; found in fishes and amphibians
ram ventilation
The production of respiratory flow in some fish in which the mouth is opened during swimming, such that water flows through the mouth and across the gills.
ovoviparity
fertilized eggs are retained within mother to complete development, but all nourishment gained from yolk sac
Oviparity
egg laying
ray
1 pelvic fin lobe, live bearing (viviparous), whip-like tail and stinging spike, absent dorsal spine