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community
an association of interacting species inhabiting some defined area
community structure
includes attributes such as the number of species, the relative abundance of species, and the kinds of species comprising a community
guild
a groups of organisms that all make their living in a similar way (seasonality, resource requirements, and life histories)
ex: seed eating animals in a desert area
life form
a combination of a plant’s structure and its growth dynamics
ex: epiphytes grow without substrate in the canapy
Preston Log-Normal Distribution
populations that demonstrate a bell-shaped/normal distribution
most species are moderately abundant
Shannon-Weiner Diversity Index
species richness: the number of species in the community
relative abundance: the species evenness of species in communities
Rank-abundance curve
portrays the relative abundance and diversity of species within a community by plotting the relative abundance of species against their rank in abundance
higher diversity has falter curves and lower diversity has steeper slope
*species are ranked by the highest species abundance (species with the most number of individuals present to the species with the lowest present)
habitat heterogeneity
complex and diverse environments allow more species diversity as it allows for more niches
mainly due to the competitive exclusion principle, which leads us to predict that coexisting species will have significantly different niches
nutrient enrichment reduces environmental complexity
as nutrient supplies increase, diversity and algae declines
It reduces the number of limiting nutrients and light becomes the single limiting resource
This means that only species that are most effective at competing for light will thrive in a community and species diversity will decline
It also reduces the need for mutualistic relationships between species; especially ectomycorrhizae
intermediate disturbance hypothesis
hypothesis that predicts that intermediate levels of disturbance promote higher levels of diversity
In lower disturbances, there is more species dominance (competition)
In higher disturbances, there is not enough recovery time for a large number of species to survive
food web
a summary of the feeding inactions within a community based on tropic levels
strong interactions in food web
the feeding activities of a few species have a dominant influence on community structure
dominant/foundation species
species that have substantial influences on community structure as a consequence of their high biomass
ex: coral on a coral reef
keystone species
relatively low biomass compared to their influence on the community structure
(1) predators might keep prey populations below their carrying capacity
(2) the potential for competitive exclusion would be low in populations kept below their carrying capacity
(3) if keystone species reduce the likelihood of competitive exclusion, their activities would increase the number of species that could coexist in communities → increase species diversity
direct interactions
between two species, including competition, predation, herbivory, and mutualism, involve positive or negative effects of one species on another without the involvement of an intermediary species
indirect interactions
one species affects through a third, intermediary species
includes trophic cascades, apparent competition, and indirect mutualism or commensalism
ex: indirect commensalism: occurs when the activity of one species indirectly — through intermediary species — benefit another species without itself being helped or harmed
apparent competition
negative impacts are the result of two species sharing a predator or herbivore or by one species facilitating populations of a predator or herbivore of the second species
Ex: if two prey species share a predator, increases in one of the prey populations may lead to increased number of the predator, which feed on and depress populations of the second prey species
primary production
production of new organic matt (biomass) by autotrophs in an ecosystem
measured as the rate of carbon uptake by primary producers or by the amount of biomass or oxygen produced
gross primary production (GPP)
total biomass produced by all autotrophs in an ecosystem
Net primary production (NPP)
biomass left over after autotrophs have met their own energetic needs
cellular respiration and maintenance of existing tissues
secondary production
the production of biomass by heterotrophic consumer organisms feeding on plants, animals, microbes, fungi, or detritus
includes consumer growth, reproduction, and mortality
terrestrial NPP
;generally limited by temperature, moisture, and nutrients
temperature: logarithmic slope because at their optimum temperature, they are most effective at photosynthesis production
precipitation: flattens because when its raining, photosynthesis cannot occur
AET: warmer and wetter climates have a higher AET and more primary production
soil nutrients: plants that have access to phosphorus and nitrogen almost double their primary production rate
actual evapotranspiration (AET)
the amount of water that evaporates and transpired off a landscape; occurs when plants open their stomata to take in CO2 and release water
increases with increased precipitation and temperature
aquatic NPP
nutrients availability is key for aquatic systems (from upwelling or breakdown of organic materials) and light availability
add nutrients —> more biomass = more chlorophyll
more chlorophyll —> higher rate of primary production
ex: highest NPP rates are found along coastlines due to nutrient runoff from land and sediment distribution
biodiversity
the genetic, philosophical, anatomical, functional, and ecosystem diversity that influences ecological processes in a community, especially in primary production
bottom up controls
influences of physical and chemical factors on an ecosystem/community
access to grazing allowed grass plants to increase their biomass as it prevents interspecific competition —> self thinning
top down controls
influences of consumers on an ecosystem/community
trophic cascade hypothesis
consumers can influence the rate of primary production in aquatic and terrestrial ecosystems
as predators increase, prey populations decrease
but the next trophic level population increases
and then the next trophic level population decreases
results in an increase, optimization, and decreases in production curve
trophic dynamics
transfer of energy from one part of an ecosystem to another
trophic level
a position in the food web and is determined by the number of transfers of energy from primary producers to that level
First level: primary producers
Second level: herbivores and detritivores
Third level: carnivores feeding on herbivores and detritivores
Fourth level: predators that feed on carnivores
ecological efficiency
the percentage of biomass produced at a lower trophic level that is transferred to biomes produced at the next trophic level, varies from 5% to 20%
Ecosystems with greater primary production generally support higher levels of secondary production
nutrient cycling
the use, transformation, movement, and reuse of nutrients in ecosystems
involves the storage of chemical elements in nutrient pools, or compartments, and the flux of nutrients between pools
nutrient pools
the amount of a particular nutrient stored in a portion, or compartment, of an ecosystem
nutrient flux
the movement of nutrients between the pools of an ecosystem
nutrient sink
a part of the biosphere where a particular nutrient is released faster than it is absorbed
phosphorus cycle
[P] marine sediments > [P] mineral deposits >>> [P] atmosphere
Phosphorus is slowly released to terrestrial and aquatic ecosystems through the weathering of rocks (apetite - type of sedimentary rock)
Terrestrial plants uptake the phosphorus
In aquatic ecosystems, phosphorus is washed into rivers and eventually makes it to the ocean where is finds ocean sediments
These are eventually transformed into phosphate-bearing sedimentary rocks that can form new land through geological uplift (tectonic plate movement)
nitrogen cycle
nitrogen sources mainly done through nitrogen fixation, lightning, and human activity
Major atmospheric pool in for of N2 available only to N-fixers
Nitrogen fixation occurs under anaerobic conditions where species oxidize sugars to obtain required energy or in high pressure and energy generated by lightning
Humans have also been able to increase agricultural productivity by rotating crops capable of nitrogen fixation and through the industrial fixation of N2 to NH3 fertilizer
once nitrogen fixation is completed, it becomes available to other organisms within the ecosystem
nitrogen can leave the organic matter pool through denitrification
carbon cycle
Carbon moves between organisms and the atmosphere as a consequence of two reciprocal biological processes: photosynthesis and respiration
Photosynthesis removes carbon dioxide from the atmosphere and respiration returns carbon dioxide to the atmosphere
In aquatic systems, CO2 must be dissolved in water before being used by autotrophs; it enters a chemical equilibrium with bicarbonate, HCO3- and carbonate CO3.
The massive burning of fossil fuels has increased the concentration of atmospheric CO2
Some C-cycles are rapid, some C may remain sequestered for long periods
terrestrial decomposition
the breakdown of organic matter accompanied by the release of carbon dioxide
significantly influenced by temperature, moisture, and the chemical decomposition of both plant litter and environment
moisture and temperature increase rates of decomposition (higher AET)
leaves with more nitrogen (less lignin) have a faster decomposition
aquatic decomposition
the more nitrogen and less lignin in the substrate, the faster the decomposition
nutrient spiraling also plays a role in decomposition rate and nutrient retention
nutrient spiraling
stream dynamics are better represented by a spiral