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Evolution (in genetic terms)
A change in allele frequencies in a population over time.
Gene pool
All the alleles of all the genes in a population.
Allele frequency
The proportion of a particular allele among all alleles of that gene in the population.
Mechanism of natural selection on allele frequencies
Individuals with advantageous alleles survive and reproduce more, so those alleles become more common.
Importance of genetic variation in evolution
Without variation, there are no different phenotypes for selection to act upon.
Three main sources of genetic variation in sexually reproducing populations
Mutation, independent assortment/crossing over in meiosis, and random fertilisation.
How mutation creates new alleles
It changes the base sequence of DNA, sometimes creating a new version of a gene.
Contribution of meiosis to genetic variation
Independent assortment and crossing over produce new combinations of existing alleles.
Contribution of random fertilisation to genetic variation
Which sperm fertilises which egg is random, combining different alleles each time.
Differential reproductive success
Differences in reproductive success between individuals in a population.
Reason some individuals produce more offspring than others
They possess phenotypes better suited to the environment, so they survive longer and reproduce more.
How differential reproductive success alters allele frequencies
Advantageous alleles are passed on more often, so their frequency increases in the gene pool.
Reason evolution lacks a predetermined direction or goal
Selection acts on existing variation and changing conditions, not towards a pre-set target.
Selection pressure
An environmental factor that affects survival and reproduction.
Examples of selection pressures
Predation, disease, competition, climate, and food availability.
Why selection pressures determine survival and reproduction of phenotypes
Individuals with traits better suited to pressures survive and leave more offspring.
Directional selection
Selection that favours individuals with phenotypes at one extreme.
Environmental conditions for directional selection
When the environment changes in a consistent direction (for example, a colder climate).
Effect of directional selection on phenotypic mean
The mean value of the trait shifts towards the favoured extreme.
Effect of directional selection on phenotypic range
Phenotypes on one side of the distribution become more common, while others become less common.
Stabilising selection
Selection that favours average phenotypes and acts against extremes.
Environmental conditions for stabilising selection
Stable, unchanging environments.
Effect of stabilising selection on mean phenotype
The mean phenotype remains roughly the same.
Effect of stabilising selection on phenotypic range
The range becomes narrower, with fewer extreme phenotypes.
Disruptive selection
Selection that favours individuals at both extremes and acts against intermediates.
Environmental conditions for disruptive selection
Environments where conditions vary in different ways within the same area or favour extremes.
Effect of disruptive selection on extreme phenotypes
Individuals at the extremes have a selective advantage and increase in frequency.
Effect of disruptive selection on intermediate phenotypes
Individuals close to the mean are at a disadvantage and decrease in frequency.
Role of disruptive selection in speciation
It can split a population into distinct groups, serving as a step towards speciation.
Species (biological species concept)
A group of similar organisms that can interbreed to produce fertile offspring.
Speciation
The formation of new species from existing ones.
Importance of reproductive isolation in speciation
Different species must be reproductively separated to remain distinct.
Role of allele frequency divergence in speciation
Differences in allele frequency lead to differences in phenotype that prevent successful interbreeding.
Allopatric speciation
Speciation that occurs after populations become geographically isolated.
Type of barrier in allopatric speciation
A physical (geographical) barrier that prevents interbreeding.
Examples of geographical barriers leading to allopatric speciation
Mountains, rivers, oceans, deserts, and ice sheets.
Why geographically separated populations experience different selection pressures
Each environment has a different climate, predators, competitors, and resources.
Action of natural selection in different environments
Different traits are favoured in different environments, so selection acts differently.
Changes in allele frequencies in isolated populations
They change in different directions, reflecting local selective pressures.
Selection of mutations in isolated populations
Different beneficial mutations arise by chance in each isolated group.
How time and continued selection result in new species in allopatry
Over many generations, accumulated differences become large enough to prevent interbreeding, forming new species.
Sympatric speciation
Speciation occurring within the same geographical area.
Mechanism of sympatric speciation without geographical isolation
Populations become reproductively isolated without physical separation.
Barriers involved in sympatric speciation
Behavioural, ecological, temporal, or mechanical barriers.
Role of behavioral changes in sympatric speciation
Changes in courtship behaviour or mate preference may prevent interbreeding.
Role of seasonal (temporal) isolation in sympatric speciation
If groups breed at different times, they do not mate with each other.
Role of polyploidy in plant sympatric speciation
A sudden increase in chromosome number can produce a new fertile plant type that is reproductively isolated from its parents.
Reproductive isolation
Barriers that prevent gene flow between populations.
Pre-zygotic isolating mechanisms
Barriers that prevent mating or fertilisation from occurring.
Examples of pre-zygotic isolating mechanisms
Different mating seasons, different courtship behaviour, mechanical incompatibility of reproductive organs, and gamete incompatibility.
Post-zygotic isolating mechanisms
Barriers that act after fertilisation.
Examples of post-zygotic isolating mechanisms
Hybrid inviability, hybrid sterility, or hybrid breakdown in later generations.
Effect of reproductive isolation on gene flow
Alleles can no longer be exchanged, allowing populations to diverge independently.
Genetic drift
Random changes in allele frequencies, especially in small populations.
Impact of genetic drift on small vs. large populations
Chance effects have a proportionally bigger impact in small populations.
Contribution of genetic drift to speciation
Drift can fix different alleles in different isolated populations, contributing to divergence.
Difference between genetic drift and natural selection
Selection is non-random and favours advantageous alleles, whereas drift is random and not necessarily adaptive.
Founder population
A small number of individuals that form a new population isolated from the original.
How the founder effect alters allele frequencies rapidly
By chance, the founder individuals may carry allele frequencies very different from the original population.
How the founder effect sets the stage for speciation
Isolation and strong drift/selection in the small population can rapidly produce a distinct gene pool.
Reason populations in different environments diverge over time
Different selection pressures, mutation events, and drift operate in each population.
Why previously identical species populations lose interbreeding ability
Accumulated genetic and behavioural differences eventually prevent successful mating or fertilisation.
Why reproductive isolation is key to defining speciation
It shows that the populations no longer share a gene pool and evolve independently.
Fossil evidence supporting evolution of new species
Fossils show changes in forms over time and the appearance of new groups not present in earlier rocks.
Punctuated equilibrium explanation of fossil record
It suggests long periods of little change interrupted by rapid speciation events, matching sudden fossil appearances.
Gradualism vs. punctuated equilibrium
Gradualism suggests slow, continuous change, whereas punctuated equilibrium suggests bursts of change separated by stasis.
Advantage of high genetic variation for future evolution
It provides a wider range of traits that may be advantageous under future environmental changes.
Why stable environments favour stabilising selection
The same conditions favour the same intermediate phenotypes, so there is little divergence.
Why changing or fragmented environments promote speciation
Changing conditions create new selection pressures in different places, promoting divergence and speciation.