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evolution
Decent with modification: the process by which species accumulate differences from their ancestors as they adapt to different environments over time; also defined as a change in the genetic composition of a population from generation to generation
fossils
a preserved remnant or impression of an organism that lived in the past
adaptation
inherited characteristic of an organism that enhances its survival and reproduction in a specific environment
natural selection
a process in which individuals that have certain inherited traits tend to survive and reproduce at higher rates than other individuals because of those traits
artificial selection
process in which humans select and breed individuals that have desired traits
homology
similarity in characteristics resulting from shared ancestry
homologous structures
structures in different species that are similar because of common ancestry; these similar structures may or may not have similar functions ex. whale fins and human arms
convergent evolution
The evolution of similar features in independent evolutionary lineages
analogous
having characteristics that are similar because of convergent evolution, not homology
biogeography
study of geographic distribution of species
phylogeny
the evolutionary history of a species or group of related species
taxon
a named taxonomic unit at ent given level of classification
phylogenetic tree
a branching diagram that represents a hypothesis about the evolutionary history of a group of organisms
branch point
The representation on a phylogenetic tree of the divergence of two or more taxa from a common ancestor. A branch point is usually shown as a dichotomy in which a branch representing the ancestral lineage splits (at the branch point) into two branches, one for each of the two descendant lineages
evolutionary lineage
the sequence of ancestral organisms leading to a particular taxon; represented by a branch in a phylogenic tree
sister taxa
groups of organisms that share an immediate common ancestor and hence are each other’s closest relatives
rooted
describing a phylogenic tree that contains a branch point representing the most recent common ancestor of all taxa in the tree
basal taxon
in a specified group of organisms, a taxon whose evolutionary lineage diverged early in the history of the group
shared ancestral character
a character that is shared by members of a particular clade but that originated in an ancestor that is not a member of that clade
shared derived character
an evolutionary novelty that is unique to a particular clade
outgroup
A species or group of species from an evolutionary lineage that is known to have diverged before the lineage that contains the group of species being studied. An outgroup is selected so that its members are closely related to the group of species being studied, but not as closely related as any study-group members are to each other
ingroup
A species or group of species whose evolutionary relationships are being examined in a given analysis
maximum parsimony
a principle that states that when considering multiple explanations for an observation, one should first investigate the simplest explication that is constant with the facts
clade
a group of species that includes an ancestral species and all of its descendants
monophyletic
Pertaining to a group of taxa that consists of a common ancestor and all of its descendants. A monophyletic taxon is equivalent to a clade.
paraphyletic
Pertaining to a group of taxa that consists of a common ancestor and some, but not all, of its descendants
polyphyletic
Pertaining to a group of taxa that includes distantly related organisms but does not include their most recent common ancestor
molecular clock
A method for estimating the time required for a given amount of evolutionary change, based on the observation that some regions of genomes evolve atconstant rates
horizontal gene transfer
the transfer of genes from one genome to another through mechanisms such as transposable elements, plasmid exchange, viral activity, and perhaps fusions of different organisms
microevolution
evolutionary change below the species level; change in the allele frequencies in a population over generations
genetic variation
differences among individuals in the composition of their genes or other DNA sequences
phenotypic plasticity
the ability of an individual genotype that is capable of producing different phenotypes when exposed to different environmental conditions
neutral variation
genetic variation that does not provide a selective advantage or disadvantage
adaptive evolution
a process in which traits that enhance survival or reproduction tend to increase in frequency in a population over time
genetic drift
a process in which chance events cause unpredictable fluctuations in allele frequencies from one generation to the next. Effects of genetic drift are most pronounced in small populations.
founder effect
genetic drift that occurs when a few individuals become isolated from a larger population and form a new population whose gene pool composition is not reflective of that of the original population
bottleneck effect
Genetic drift that occurs when the size of a population is reduced, as by a natural disaster or human actions. Typically, the surviving population is no longer genetically representative of the original population
gene flow
the transfer of alleles from one population to another, resulting from the movement of fertile individuals of their gametes
relative fitness
the contribution an individual makes to the gene pool of the next generation , relative to the contribution of other individuals in the population
directional selection
Natural selection in which individuals at one end of the phenotypic range survive or reproduce more successfully than do other individuals
disruptive selection
Natural selection in which individuals on both extremes of a phenotypic range survive or reproduce more successfully than do individuals with intermediate phenotypes
balancing selection
neutral selection that maintains two or more phenotypic forms in a population
heterozygote advantage
grater reproductive success of heterozygous individuals compared with homozygotes; tends to preserve variation in a gene pool
gene pool
The aggregate of all copies of every type of allele at all loci in every individual in a population. The term is also used in a more restricted sense as the aggregate of alleles for just one or a few loci in a population
population
a group of individuals of the same species that live in the same area and interbreed, producing fertile offspring
Hardy-Weinberg Equilibrium
The state of a population in which frequencies of alleles and genotypes remain constant from generation to generation, provided that only Mendelian segregation and recombination of alleles are at work, null hypothesis
allopathic speciation
the formation of new species in populations that are geographically isolated from one another
biological species concept
definition of a species as a group of populations whose members have the potential to interbreed in nature and produce viable, fertile offspring, but do not produce viable, fertile offspring with members of other such groups
ecological species concept
definition of a species in terms of ecological niche, the sum of how members of the species interact with the nonliving and living parts of their environment
hybrids
offspring that results from the mating of individuals from two different species or from two true-breeding varieties of the same species
hybrid zone
A geographic region in which members of individuals from two different species meet and mate, producing at least some offspring of mixed ancestry
macroevolution
Evolutionary change above the species level. Examples of macroevolutionary change include the origin of a new group of organisms through a series of speciation events and the impact of mass extinction on the diversity of life and its subsequent recovery.
microevolution
Evolutionary change below the species level: change in the allele frequencies in a population over generations
morphological species concept
definition of a species in terms of measurable anatomical criteria
postzygotic barriers
a reproductive barrier that prevents hybrid zygotes produced by two different species from developing into viable, fertile adults
prezygotic barriers
a reproductive barrier that impedes mating between species or hinders fertilization if interspecific mating is attempted
punctuated equilibria
In the fossil record, long periods of apparent stasis, in which a species undergoes little or no morphological change, interrupted by relatively brief periods of sudden change.
reproductive isolation
the existence of biological factors (barriers) that impede members of two species from producing viable, fertile offspring
speciation
an evolutionary process in which one species splits into two or more species
adaptive radiation
Period of evolutionary change in which groups of organisms form many new species whose adaptations allow them to fill different ecological roles in their communities
mass extinction
The elimination of a large number of species throughout Earth, the result of global environmental changes
Pangaea
the supercontinent that formed near the end of the Paleozoic era, when plate movements brought all the landmasses of life together
plate tectonics
the theory that the continents are part of great plates of Earth’s crust that float on hot, unyielding portion of the mantle. Movements in the mantle cause the continents to move slowly over time
radiometric dating
common techniques used to determine the age of a fossil, based on the decay of radioactive isotopes
heterochrony
evolutionary change in the timing or rate of an organism’s development
paedomorphisis
the retention in an adult organism of the juvenile features of its evolutionary ancestors
homeotic genes
Any of the master regulatory genes that control placement and spatial organization of body parts in animals, plants, and fungi by controlling the developmental fate of groups of cells
prokaryotes
an organism that has a prokaryotic cell: an informal term for an organism in either domain Bacteria or domain Archea
protocells
an abiotic precursor of a living cell that had a membrane-like structure that maintained an internal chemistry different form that of its surroundings
pathogens
an organism or virus that causes disease
ribozymes
an RNA molecule that functions as an enzyme, such as an intron that catalyzes its own removal during RNA splicing
nucleoid
A non-membrane-enclosed region in a prokaryotic cell where its chromosomes is located
plasmids
A bacterial plasmid carrying genes that confer resistance to certain antibiotics.
transformation
The process by which a cell in culture acquires the ability to divide indefinitely, similar to the division of cancer cells. (2) A change in genotype and phenotype due to the assimilation of external DNA by a cell. When the external DNA is from a member of a different species, transformation results in horizontal gene transfer
transduction
A process in which phages (viruses) carry bacterial DNA from one bacterial cell to another. When these two cells are members of different species, transduction results in horizontal gene transfer. See also signal transduction pathway
conjugation
In prokaryotes, the direct transfer of DNA between two cells that are temporarily joined. When the two cells are members of different species, conjugation results in horizontal gene transfer
protists
An informal term applied to any eukaryote that is not a plant, animal, or fungus. Most protists are unicellular, though some are colonial or multicellular
endosymbiosis theory
The theory that mitochondria and plastids, including chloroplasts, originated as prokaryotic cells engulfed by host cells. The engulfed cell and its host cell then evolved into a single organism
endosymbiosis
A mutually beneficial relationship between two species in which one organism lives inside the cell or cells of another organism
secondary endosymbiosis
A process in eukaryotic evolution in which a heterotrophic eukaryotic cell engulfed a photosynthetic eukaryotic cell, which survived in a symbiotic relationship inside the heterotrophic cell