Evolution
A change in heritable traits within a population, occurs via natural selection
Lamarckism
previous evolutionary (false), said evolution occurred via acquired traits during an organism's lifetime
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Evolution
A change in heritable traits within a population, occurs via natural selection
Lamarckism
previous evolutionary (false), said evolution occurred via acquired traits during an organism's lifetime
Sequential Evidence
Evolutionary change visible in DNA/RNA/AA, fewer differences=more closely related
Sequence differences accounted by the splitting of ancestral species
Gene Family
Ex. Hox Gene, determines body plan during development, found in organisms w/ clear head-to-tale axis
Selective breeding
Artificial selection, deliberate breeding process to produce desired traits (starts with one species)
Purpose: increase frequency of desired traits in short period of time
Mimics and provides evidence of evolution
Diversity in domesticated animals & Plants (2)
in between breeds and from original wild species
Homologous Structures
Similar anatomical structures w/ different functions, inherited from common ancestors but adapted for different functions
ex. Pentadactyl limbs
Pentadactyl limbs
Similar bone arrangement of five-fingered limb shared in mammals, birds, amphibians, and reptiles
However each limb shows adaptation to their mode of locomotion
Analogous Structures
Different anatomical structures that have similar functions, result of convergent evolution
Inherited from different ancestor, but lived in similar habitats w/ similar selective pressures, thus similar adaptations
Speciation
Creation of new species from pre-existing species
Two separate species begin to evolve independently and can no longer interbreed when brought back together (biological species concept)
Reproductive Isolation
When barriers (not always physical) prevent gene flow between gene pools of two populations
ex. geographic separation (physical barriers)
Divergent/Differential Selection
Different selective pressures cause a population to become different over time
Differences:
-climate
-predation
-competition
Bonobos & Chimpanzees
Separated by Congo river, water level lowered at one point and chimps crossed to Bonobo side
Chimps then subject to same selective pressures as Bonobos and become absorbed into bonobos
Types of Speciation
Allopatric Speciation & Sympatric Speciation
Both require reproductive isolation:
Allopatric Speciation
Geographic barrier that creates physical isolation
Sympatric Speciation (3 types)
The formation of new species in populations that live in the same geographic area
-Behavioral isolation
-Temporal Isolation
-Mechanical isolation
Behavioral Isolation
two populations have different courtship rituals or mating rituals
Temporal Isolation
Two populations breed during different times of the year
Mechanical Isolation
Two populations are anatomically incompatible and cannot transfer sperm
Adaptations
Heritable traits that allow individuals to be better suited to their environments
Adaptive Radiation
Pattern of diversification where species from a common ancestor occupy different, vacant niches (ecological roles) to minimize competition and increase biodiversity
Darwin's Finches
Rapid evolution of single species, results in diversification as individuals with adaptations that match their unique selective pressures survive and reproduce
Seen in beak shape
Finches eventually occupy different niches
Interspecific Hybrids
result of cross-breeding different species, combines useful traits of both (hybrid vigor), but usually sterile
Barriers to hybridization
In evolutionary terms, energy spent on producing sterile offspring is a waste, thus, organisms use techniques like courtship rituals to prevent it
Polyploidy
Organism has more than 2 sets of homologous chromosomes, chromosome duplicate but fail to divide in meiosis
Autotetraploid
(4n), all chromosomes come from same organism, gametes were diploid
When fuse w/ haploid gamete, offspring are triploid (3n) and are sterile
Therefore autotetraploids = different species (biological species concept)
Hybrid
When two species cross breed, contains one set of chromosomes from each parent, chromosomes do not form homologous pairs and hybrid is sterile
Allotetraploid
(4n) - chromosomes are from different species
occurs when chromosomes duplicate but don't divide in meiosis
Can interbreed with other allotetraploids but not parents (therefore different species)
Smartweed
Example of alloploidy, two diploid gametes of two different species mated and formed tetraploid smartweed
Why is polyploidy more common in plants?
Self-pollination - plants possess both male and female parts (therefore don't need to worry about 1 male and 1 female polyploid plant for reproduction)
Asexual reproduction - Infertile polyploids can reproduce asexually
Advantages of Polyploid Crops
Produce seedless (infertile) fruits
Shows hybrid vigor by growing larger, showing better disease resistance
Asexual Reproduction (4)
1. involves one parent
2. uses mitosis
3. offspring are genetically identical to each other and to parent (no genetic variation)
4. adapted to stable environment
Asexual Reproduction Examples (3)
1. binary fission
2. budding
3. vegetative propagation
Asexual Reproduction Advantage
produces genetically identical offspring by individuals that are adapted to existing environment
also rapid reproduction (no need for mate), requires less energy
Sexual Reproduction (4)
1. involves two parents (male and female)
2. involves meiosis and fertilization
3. offspring are genetically unique from each other and parents (genetic variation)
4. may be better adapted to changing environment
Sexual Reproduction Examples (3)
internal fertilization, external fertilization, pollination
Sexual Reproduction Advantage
produces offspring with variation which increases adaptability to environmental changes
Meiosis
produces unique gametes by mixing parental genotypes, variation occurs via crossing over and independent assortment
Fertilization
combines gametes to produce original offspring different from parents
-it doubles chromosome number, requiring meiosis to halve it
-gametes are haploid, zygotes are diploid
Reproductive System
organ system responsible for creating offspring and passing on genetic information
-unlike other systems, structures differ by gender
Testis (pl. testes)
produces sperm and testosterone
Epididymis
stores sperm until it matures (gains mobility) and is ejaculated
Scrotum
skin fold that holds testes at a lower body temp (about 2 C colder)
-allows sperm to survive
Vas Deferens
muscular duct that propels sperm during ejaculation
Seminal Vesicle and Prostate Gland
secretes fluids with nutrients, prostaglandin, and anticoagulating enzymes to form semen
Prostaglandin
causes uterine contractions
Bulbourethral Gland
secretes alkaline fluid that neutralizes acidity in urine
Penis
enters vagina for semen ejaculation
Urethra
transfers semen and urine out of body via penis
Ovary
produces egg and oestradiol (type of estrogen)/progesterone
Fimbria (pl. fimbriae)
tissue that sweeps released oocyte (egg) into oviduct
Oviduct/Fallopian Tube
carries oocyte/embryo to uterus
-site of fertilization
Uterus
muscular organ where embryo develops
Endometrium
inner lining of uterus
-thickens in preparation of implantation of embryo and lost during menstruation
Cervix
neck of uterus
-protects fetus during pregnancy, dilates for childbirth
Vagina
birth canal
Vulva
outer area that protects internal reproductive organs
Puberty
biological transition from childhood to adulthood marked by secondary sexual characteristics and reproductive maturity
-regulated by hypothalamus-pituitary-gonadal (HPG) axis
Hypothalamus
secretes increasing amounts of gonadotropin-releasing hormone (GnRH) in childhood
-stimulates anterior pituitary (front) to release gonadotropins
like luteinizing hormone (LH) and follicle-stimulating hormone (FSH)
*part of puberty*
Female Sex Hormones
estrogen (oestradiol) and progesterone, both from ovaries
-causes development of secondary sex characteristics (body hair, breast development)
-regulates menstrual cycle
Male Sex Hormones
Testosterone (from testes)
-causes development of secondary sex characteristics (facial/body hair, muscle mass, deep voice)
-growth of testes/penis
-initiation of sperm production
Menstrual Cycle
produces an ovum (egg) which allows pregnancy
-begins at puberty, ends at menopause
-lasts about 28 days
Ovarian Cycle
The 28 days of the menstrual cycle as they apply to events in the ovary. The ovarian cycle has three subphases: the follicular phase, ovulation, and the luteal phase.
Follicular Phase (days 1-14)
Purpose: growth and maturation of follicles
-FSH stimulates follicle growth --> release oestradiol
-oestradiol release more FSH and begins LH (positive feedback, process repeats w/ more FSH—> more oestradiol)
*oestradiol inhibits FSH at higher levels (not yet)*
Follicles
clusters of cells surrounding a single egg
Ovulation (day 14)
Purpose: release oocyte from follicle
-LH causes surge in ovulation
-positive feedback w/ FSH/LH ends when follicle ruptures
*LH then inhibits FSH (negative feedback*
Luteal Phase (day 15-28)
Purpose: LH converts ruptured follicle to corpus luteum
-progesterone thickens and maintains endometrium
-with oestradiol, it inhibits both FSH and LH (negative feedback)
-decrease in FSH leads to decrease oestradiol (just making less, not gone)
oestradiol also supports uterine development
Corpus Luteum
helps produce progesterone
Uterine Cycle
The cycle that prepares the uterus for the growth of an embryo
3 Phases:
Menstrual Phase
Proliferative Phase
Secretory Phase
Menstrual Phase (day 1-5)
Purpose: drop in progesterone and estrogen sheds endometrium
-if fertilization does not occur, corpus luteum is converted to corpus albicans
No corpus luteum leads to decrease progesterone leads to increase FSH which begins cycle
Proliferative Phase (6-14)
Purpose: regrow endometrial lining
-if fertilization occurs, embryo releases hormones to sustain the corpus luteum
corpus luteum makes progesterone, which thickens/maintains endometrium for pregnancy
Secretory Phase (day 15-28)
Purpose: endometrium prepares for implantation
-increase blood supply and glandular secretions
-progesterone maintains uterine lining
Gametogenesis
process in which diploid cells become haploid gametes
-in males, sperm is produced via spermatogenesis
-in females, egg is produced via oogenesis
*both processes involve MITOSIS, MEIOSIS, and DIFFERENTIATION*
Spermatogenesis
production of spermatozoa (sperm)
location: seminiferous tubules of testes
Spermatogenesis Process (5)
CONTINUOUS
1. during puberty, spermatogonia (2n) continuously undergo mitosis
2. cells that divide from spermatogonia grow/mature into primary spermatocytes (2n)
3. after meiosis I, forms two secondary spermatocytes (n)
3. after meiosis II, forms four spermatids (n)
4. during differentiation, acquires a tail and becomes spermatozoa
5. detach from sertoli cells and move out of testes
Spermatogonia
2n
sperm stem cell, continuously divides but never differentiated (divides to produce primary spermatocytes)
Primary Spermatocytes
2n
produced by division of spermatogonia, undergo meiosis I
Secondary Spermatocytes
n
2 produced by meiosis I of primary spermatocytes, undergo meiosis II
Spermatids
n
4 produced by meiosis II of secondary spermatocytes, consists of the "head" of the sperm
Spermatozoa
produced by differentiation of spermatids, gets tail
Sertoli Cells
mother cells that provide nutrients for spermatids
Oogenesis
production of ova (egg)
location: ovaries
*DISCONTINUOUS*
Oogenesis Process Before Birth (2)
1. before birth, oogonia (2n) continuously undergoes mitosis
2. cells that divide from oogonia grow/mature into primary oocytes, enters prophase I (meiosis) and pauses until puberty
Oogenesis Process During Puberty (3)
1. during puberty, one primary oocyte completes meiosis I during each menstrual cycle
2. forms one larger secondary oocyte (n) which gets all the cytoplasm, and enters meiosis II & pauses until fertilization
also forms one smaller polar body that eventually degenerates due to unequal cytoplasmic division
Oogenesis Process During Ovulation
1. secondary oocyte is released from follicle and enters oviduct (fallopian tubes)
2. if fertilized, meiosis II is completed
3. forms ovum and another polar body via unequal division of cytoplasm
*if not fertilized, secondary oocyte is just removed from body w/ rest of period*
Oogonia
stem cell that produces primary oocytes in mitosis, female version of spermatogonia
Primary Oocyte
produced by division of oogonia, enters prophase I before birth and pauses until puberty
Secondary Oocyte
formed when meiosis I is completed during each menstrual cycle, enters meiosis II and pauses until fertilization
Ovum
fertilized egg, formed if secondary oocyte is fertilized and completes meiosis II after ovulation
Male/Female Gamete Production
Male: millions of sperm (small, motile)
-produced in testes via spermatogenesis
-released continuously
Female: limited number of ova (large, sessile)
-produced in ovaries via oogenesis
-released cyclically
Male Gamete Structure/Adaptations (4)
1. sperm streamlined for motility
2. head - haploid nucleus
3. midpiece - large amounts of mitochondria to fuel movement via atp
4. tail - flagellum for motility, made of microtubules
Sperm Head
contains acrosome, paired centrioles, and has minimal cytoplasm
Acrosome
has enzymes to degrade jelly coat of egg
Paired Centrioles
needed by zygote to begin mitosis
Female Gamete Structure/Adaptations (4)
1. egg contains large amount of cytoplasm w/ nutrients
2. zona pellucida
3. corona radiata
4. cortical granules
Zona Pellucida
protective jelly coat
Corona Radiata
follicular layer that provides structural support
Cortical Granules
released during fertilization to prevent polyspermy
Male vs. Female Hormone Regulation
Male: testosterone regulates spermatogenesis and secondary sex characteristics (muscle, body hair)
Female: estrogen/progesterone regulate ovarian and menstrual cycles and secondary sexual characteristics (breast development)
Male vs. Female Reproductive Roles
Male: produce/deliver sperm for fertilization
Female: produce eggs, provide site for fertilization, support embryo development