variation, inheritance, differential reproductive success
what are the components of natural selection
adaptions
characteristics that enhance survival or reproduction of organisms compared to alternate character states
natural selection
What accounts for the evolution of adaptions?
life history strategy
-The schedule and manor in which an organism invests in survivorship and reproduction over lifespan -reveals trade-offs
natural selection
-the nonrandom survival of randomly varying replicators -any consistent difference in fitness among different classes of biological entities
no
Is natural selection goal-directed?
fitness
biological entities avg per capita rate of increase in numbers
probability of survival to reproductive age, avg # of offspring produced via female and male functions
What are key components of fitness?
sexual selection
special kind of natural selection that results from the competition for mates.
variation in fitness, external force, reproductive success
Natural selection exists whenever there is _______. Natural selection is not an ________. It is a name for a statistical difference in _________ among genes, organisms, (and populations)
selection for
describes trait that creates advantage/higher fitness
selection of
any trait correlated with advantageous trait but has no effect on fitness, piggybacks
selection of
In the ball example, would red balls be selection for or of?
male/male competition and female choice
What are the 2 types of sexual selection?
stronger
When some features may be subjected to conflicting selection pressures (e.g. sexual selection and predation), the direction of evolution depends upon which is ________.
selfish genetic elements
In many species, there exist ____________, which are transmitted at a higher rate than the rest of an individual’s genome and are detrimental.
segregation distortion
Change 1:1 ratio of being in gamete towards certain trait Ex: allele has 90% frequency in sperm
group selection
organisms do things for good of species
individuals, groups, self-sacrificing
Problems with group selection.... -The survival of alternative alleles is much more likely to be determined by differences in the reproductive success of genetically different _________ than by genetically different _________. -Behaviors that appear to be ________ often really aren’t
females who raised extra offspring
Who would benefit more: females that produced fewer chicks for the benefit of the species or those females who raised extra offspring?
altruism
one individual aids another in a way that improves others fitness but declines their own as consequence
kin selection
Seemingly altruistic behaviors evolve by ________
are related to individual performing behavior
An allele for altruistic behavior can increase in frequency if the beneficiaries of the behavior ________.
gene, survival in future generations
gene for altruism leads to aid of others with same _____, therefore increasing fitness of that and leading to its ________.
misfirings of kinship
Loss of offspring-> hormonally inclined to care
co-options
-features evolved in the past under different conditions and serve new functions -Ie: beaks good for fruits and nuts also break through sheep skin
no
Are all traits of organisms adaptations?
physics or chemistry, genetic drift, correlated with other adaptive traits, phylogenetic history
Not all traits are adaptations... 1.Traits may be consequences of _______ 2. Traits may have evolved by ________. 3. Traits may have evolved because they were _________. 4. Traits may be consequences of ___________.
complexity, appearance of design, experiments, the comparative method
what can be used to infer adaptation?
The necessity of adaptation
Perfection
Progress
Harmony and the balance of nature
Morality and ethics
what should not be expected of natural selection and adaption
Variation
Pleiotropic effects
Gene flow
No foresight
Physics
what are some constraints of natural selection?
no
Is natural selection the same as evolution?
Natural selection is a mechanism that allows individuals best suited to an environment to pass on genes. Evolution by natural selection is the change in acquired traits over generations because of this.
Why is natural selection different than evolution by natural selection?
no
Does natural selection act at the same rate on all loci in a genome
positive
A feature cannot evolve by natural selection unless in makes ______ contribution to reproduction or survival
fitness, genotype, fitness and genotype
Consequences of natural selection depend on… -Relationship between phenotype and _______ -Relationship between phenotype and _________ -So relationship between ___________ determines whether or not evolutionary change occurs
stabilizing selection
-Selection where average does the best (reduced fitness at extremes) -Reduces varience -In environments stable for long periods of time
directional
-selection seen in environments w/ environmental change -Favors one more extreme value compared to the mean
diversifying selection
-Individuals w/ mean trait do worst -Extreme traits do better than mean (not necessarily equally) -Common w/ environmental change or move to new area
fixed
As long as no other evolutionary factors intervene, a character state with any advantage will be _______ by natural selection.
fitness
average lifetime contribution of individuals of genotype to the population after 1 or more generations
Absolute Fitness (R )
Per capita growth rate
relative fitness
Genotypes value of R relative to some other genotype
Mean fitness (w)
Average fitness of individuals in population relative to the fittest genotype
proportion of individuals surviving to each age class
fecundity of each age class
age of reproduction
What does R depend upon?
Recurrent mutation and gene flow
Why do deleterious alleles persist?
frequency dependent selection
The fitness of a genotype is dependent upon its frequency in a population
inverse frequency dependent selection
High fitness bc a gene is rare Fitness declines when fitness increases
Positive frequency dependent selection
Fitness increases as a gene becomes more common
1:1
What is the approximate sex ratio?
chance or randomness
physical causes can result in any one of several outcomes, we cannot predict what the outcome will be
chance and nonrandom
Most phenomena are affected by both ______ and ______ factors
nonrandom
Adaptations evolve by natural selection which is ________
genetic drift and natural selection
What are the two most important random causes of changes in allele frequencies in populations
genetic drift
The process of random fluctuation in allele frequencies due to sampling effects in finite populations.
fluctuate over time, heterozygotes, diverge
In genetic drift...
In a finite population, allele frequencies _______, even in the absence of natural selection.
Some alleles are fixed, others are lost, and the fraction of __________ in the population decreases over time.
Separate populations _______ in their allele frequencies and in terms of which alleles are present.
likely, not be included in genes that form zygotes, sample, sampling error
Genetic drift as sampling error.... -If an allele is rare, it is _______ to be lost by chance alone. -One or a few copies of the allele may_______. -Genes in one generation are a _______ of the genes in the previous generation. -Any sample is subject to random variation or ________.
stabilizing force
There is no _______ so allele frequencies do not stay the same and instead drift to either 0 or 1
may vary
allele frequencies ______ among the demes
fixed, heterozygotes, frequency, diverge, small
Evolution by genetic drift...
Allele frequencies fluctuate at random in a population and eventually one or another allele becomes ____.
As the frequency of one allele approaches 1, the frequency of _______declines.
An allele’s probability of fixation equals its _______ at that time.
Populations with the same initial allele frequency ________, and a proportion of the populations is expected to become fixed for that allele.
Evolution by genetic drift proceeds faster in ________ populations.
effective population size
The number of individuals that contribute genes to the next generation
greater
Due to effective population size, the rate of genetic drift will be _____ than expected from populations census size
founder effects
Effects of genetic drift that occur with reduction in population size
bottlenecks
-Sudden restrictions in population size -Genetic drift has more pronounced effect on smaller population
selectively neutral, adaptive
The neutral theory of molecular evolution states... -Most of genes organisms carry _________ -Most genetic variation lacks _________ significance
no
Does the neutral theory hold that the morphology, physiology, and behavior of organisms evolves by random genetic drift?
natural selection
In the case of an environmental difference does natural selection or evolution occur?
Inverse frequency dependent selection
what keeps the sex ratio around 1:1?