Evolution

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Last updated 1:55 PM on 9/2/26
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113 Terms

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how and when the earth formed

  • 4.5 billion years ago

  • as mass of developing planet increased, heat generated by force of gravity and radioactive decay caused interior to melt

  • produced dense metallic core surrounded by cooler, semi-solid mantle

  • outside of that was crust - solidified to form continents and seafloor


<ul><li><p>4.5 billion years ago</p></li><li><p>as mass of developing planet increased, heat generated by force of gravity and radioactive decay caused interior to melt</p></li><li><p>produced dense metallic core surrounded by cooler, semi-solid mantle</p></li><li><p>outside of that was crust - solidified to form continents and seafloor</p></li></ul><p></p>
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conditions of early earth (and how it was formed)

  • cooling of the crust caused gases from the hot interior to escape through volcanoes

  • atmosphere contained hydrogen, water vapour, methane, ammonia, hydrogen sulfide

  • lacked oxygen


<ul><li><p>cooling of the crust caused gases from the hot interior to escape through volcanoes</p></li><li><p>atmosphere contained hydrogen, water vapour, methane, ammonia, hydrogen sulfide</p></li><li><p>lacked oxygen</p></li></ul><p></p>
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cyanobacteria/stromatolite importance and overview

  • cyanobacteria in rocks - oldest widely accepted fossils

  • formed stromatolites - layers produced as CaCO3 precipitated over growing bacterial filaments

  • released oxygen into the atmosphere by photosynthesis


<ul><li><p>cyanobacteria in rocks - oldest widely accepted fossils</p></li><li><p>formed stromatolites - layers produced as CaCO3 precipitated over growing bacterial filaments</p></li><li><p>released oxygen into the atmosphere by photosynthesis </p></li></ul><p></p>
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processes necessary for life to begin on earth

  • synthesis of simple organic molecules (e.g. amino acids) and ultimately synthesis of larger organic polymers (e.g. proteins)

  • origin of molecules that can self-replicate

    • only molecules known to do this are RNA and DNA

  • packaging of self-replicating molecules into membranes with internal environment different in composition from their surroundings. closed membrane vesicles form spontaneosuly from lipids, can maintian diff chemical compositions between intrace and extracellular environments


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primordial soup theory

  • life on earth started in ancient ocean/pond from mixture of simple chemicals

  • lightning hit chemicals to form amino acids

  • miller-urey experiment used to see if possible for simple inorganic substances to react and form complex organic molecules

  • used water, methane, ammonia, hydrogen

  • found 13/20 amino acids formed, high conc of adenine, 15% of carbon compounds were organic


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meteor theory

  • earth subjected to lengthy barrage of meteor showers and emissions from volcanoes

  • caused atmosphere to contain precursors needed to make organic molecules


<ul><li><p>earth subjected to lengthy barrage of meteor showers and emissions from volcanoes </p></li><li><p>caused atmosphere to contain precursors needed to make organic molecules</p></li></ul><p></p>
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deep sea/hydrothermal vent theory

  • vents warmed ocean water to temperatures conducive to life processes

  • released hydrogen rich molecules and minerals to the water that early organisms could use for energy


<ul><li><p>vents warmed ocean water to temperatures conducive to life processes</p></li><li><p>released hydrogen rich molecules and minerals to the water that early organisms could use for energy</p></li></ul><p></p>
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to be considered living, an organism must…

  • consist of cells

  • control internal environment

  • control exchange of materials

  • remove waste

  • respond to stimuli

  • use energy

  • grow and reproduce


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ribozyme

  • RNA molecule that can speed up specific chemical reactions in body

  • self replicating

  • catalyse range of reactions

  • ability of RNA to be double helix or single strand allow them to self replicate and replicate other RNA molecules


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RNA world hypothesis

  • theory began due to discovery of ribozymes

  • early life used RNA not DNA - first cells consisted of RNA molecule enclosed in vesicle (protocells)

  • theorised because ribozymes self replicated and performed current role of proteins cayalyzing reactions


<ul><li><p>theory began due to discovery of ribozymes</p></li><li><p>early life used RNA not DNA - first cells consisted of RNA molecule enclosed in vesicle (protocells)</p></li><li><p>theorised because ribozymes self replicated and performed current role of proteins cayalyzing reactions</p></li></ul><p></p>
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assumptions made in RNA world hypothesis

  • genetic information was transmitted and stored by RNA in first cells

  • same replication of RNA in first cells as modern

  • metabolic reactions catalysed by RNA rather than enzymes in first cells

  • RNA molecules with ability to catalyse reactions under certain circumstances - ribozymes


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assumptions against primordial soup theory

  • geochemists now believe that early atmosphere did not contain same proportions of gases as in miller-urey

  • water breaks down protein chaincs in amino acids (or other constituents) - makes it v difficult to produce proteins/other polymers in soup

  • amount of energy required is greater than likely energy generated by electrical storms


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LUCA

  • Last Universal Common Ancestor

  • not a ref for first life on earth but ref to population from which all existing life derived from


<ul><li><p>Last Universal Common Ancestor </p></li><li><p>not a ref for first life on earth but ref to population from which all existing life derived from </p></li></ul><p></p>
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prokaryotes existed before eukaryotes evidence - fossil record


  • first cells existed 3.5 billion yrs ago - prokaryotes that didnt need O2 - cyanobacteria in stromatolites

  • 2.3 billion yrs ago - photosynthetic prokaryotes - changed atmopsheric conditions by adding 02

  • oldest eukaryotic fossil found 2.3 billion yrs ago - unicellular algae

  • earliest multicellular eukaryotic fossil 1.6 billion yrs ago - red algae


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prokaryotes existed before eukaryotes evidence - structural features

knowt flashcard image
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endosymbiotic theory

  • proposes chloroplasts and mitochondria were once ‘free-living’ prokaryotic organisms that were engulfed by another cells by endocytosis

  • they evolved within host cell in symbiotic relationship

    • host took prokaryotic cell that may have capacity to photosynthesise (hence supply organic molecules) or perform aerobic respiration (provide energy for cells)

    • engulfed prokaryotes benefited by obtaining shelter and protection from predators

  • other endosymbiotic events occured during evolution of eukaryotes (e.g. infolding of cell membrane to make ER and nucleus)


<ul><li><p>proposes chloroplasts and mitochondria were once ‘free-living’ prokaryotic organisms that were engulfed by another cells by endocytosis </p></li><li><p>they evolved within host cell in symbiotic relationship</p><ul><li><p>host took prokaryotic cell that may have capacity to photosynthesise (hence supply organic molecules) or perform aerobic respiration (provide energy for cells)</p></li><li><p>engulfed prokaryotes benefited by obtaining shelter and protection from predators </p></li></ul></li><li><p>other endosymbiotic events occured during evolution of eukaryotes (e.g. infolding of cell membrane to make ER and nucleus)</p></li></ul><p></p>
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evidence for endosymbiotic theory

facts that both mitochondria and chloroplasts:

  • have inner membranes resembling cell memrbranes of prokaryotes more than cell membrane of eukaryote they are found in

  • divide by binary fission - new cant be produced as directed by genes in nucleus

    • however rely on nuclear proteins for division

  • have circular DNA

  • have own protein synthesis system (inc. ribosomes that are incidentally more similar in size and structure to prokaryotic than eukaryotic)


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evidence for common ancestry of all living things

  • universal presence of DNA

  • suggests that early prokaryotes evolved on their own first, then eukaryotes appeared biollions of yrs later (approx 2.7)


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mutations (general) - inhertiance, define, types

  • permanent change in sequence of nucleotides of DNA

  • random changes to DNA sequence of an organism - can be beneficial, harmful, or no effect

  • sources of new forms of particular gene s(alleles) in population of species

  • somatic cell mutations are not inherited, mutations in gametes or germline cells are

  • are generally pt mutation to single gene, may or may not affect one protein only


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mutations - evolution

  • ultimate source of genetic variation in species

  • if beneficial → adaptation, if harmful → genetic disease

  • driving force of evolution - phenotypical features arise from them; beneficial mutations provide individuals with a greater survival chance and are passed through generations and become more abundant

  • sources of new forms of particular gene s(alleles) in population of species

  • accumulation of pt mutations to diff genes in diff populations, over long time period, contributes to array of differences between species


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affect of population seperation on mutations

  • If separated for long time, accumulate different mutations in DNA

  • larger time separated, the more likely there is a greater difference


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molecular clocks

  • compares evolution of biological molecules (e.g., DNA and proteins) to estimate time in history when species diverged from a common ancestor

  • differences in aa sequences for specific proteins in species are proportional to time since species diverged from CA

  • all proteins have diff rates of change but occur at constant rate

  • if constant rate is true for all species, diff in aa sequence between 2 species can be used to determine approx time of divergence

  • molecular clock must be calibrated properly

    • generally use fossil record, geological evidence and species samples

    • more samples for clock, more precise calibration


<ul><li><p>compares evolution of biological molecules (e.g., DNA and proteins) to estimate time in history when species diverged from a common ancestor </p></li><li><p>differences in aa sequences for specific proteins in species are proportional to time since species diverged from CA</p></li><li><p>all proteins have diff rates of change but occur at constant rate </p></li><li><p>if constant rate is true for all species, diff in aa sequence between 2 species can be used to determine approx time of divergence</p></li><li><p>molecular clock must be calibrated properly</p><ul><li><p>generally use fossil record, geological evidence and species samples</p></li><li><p>more samples for clock, more precise calibration</p></li></ul></li></ul><p></p>
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phylogenics - definition and principle behind analysis

  • phylogenics - finding evolutionary relationships among organisms

  • DNA, RNA, and protein sequences analysed - greater similarity, fewer mutations required to convert one sequence to other ∴ more recently share common ancestor


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phylogenetic tree

  • common ancestor occurs at the ‘root’ of the tree

  • each branching pt (node) indicates where species diverged

  • longest separation means more differences through mutation (more time that mutations have to accumulate in DNA sequence


<ul><li><p>common ancestor occurs at the ‘root’ of the tree</p></li><li><p>each branching pt (node) indicates where species diverged </p></li><li><p>longest separation means more differences through mutation (more time that mutations have to accumulate in DNA sequence</p></li></ul><p></p>
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comparative genomics

  • comparing complete genomes, genes or aa sequences of different species using a variety of tools

  • requires use of computer-based analysis to locate differences in sequences and regions of similarity

  • e.g. of program ‘Clustal Omega’ - global multiple sequence alignment program for DNA, RNA, proteins


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benefits of comparative genomics

  • major contributor to understanding organisms (inc. the human genome)

  • Important in identifying genes that are essential to life and contributing to understanding of how gene signals control gene function across a range of species

  • powerful tool for studying evolution and evolutionary relatedness between organisms and humans

  • wide range of applications in agriculture and biology


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DNA sequencing

  • used to work out the sequence of bases in a specific gene/genome

  • sample of DNA from species can be sequenced to compare evolutionary relationships

  • more similar genetic code, more closely related species - less time for mutations to accumulate

  • similarity therefore also indicates recent separation from common ancestor between species

  • feasible using comparative genomics software

    • construct phylogenetic tree using DNA sequences

    • Clustal Omega produces multiple sequence alignments of divergent sequences, calculates best match for selected sequences


<ul><li><p>used to work out the sequence of bases in a specific gene/genome</p></li><li><p>sample of DNA from species can be sequenced to compare evolutionary relationships</p></li><li><p>more similar genetic code, more closely related species - less time for mutations to accumulate</p></li><li><p>similarity therefore also indicates recent separation from common ancestor between species</p></li><li><p>feasible using comparative genomics software</p><ul><li><p>construct phylogenetic tree using DNA sequences </p></li><li><p>Clustal Omega produces multiple sequence alignments of divergent sequences, calculates best match for selected sequences </p></li></ul></li></ul><p></p>
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rRNA sequencing fixxx

  • rRNA is component of ribosomes - found in all liing organisms

  • prokaryotes - 16s rRNA gene codes for small ribosomal subunit

  • eukaryotes - 18s rRNA gene codes for small ribosomal subunit

  • cloesly related species (same genus) have similar rRNA sequences

    • info can be used to make phylogenetic trees using computer-based software

  • can also be used as qualititative method to determine unknown species and diff types in sample


<ul><li><p>rRNA is component of ribosomes - found in all liing organisms </p></li><li><p>prokaryotes - 16s rRNA gene codes for small ribosomal subunit</p></li><li><p>eukaryotes - 18s rRNA gene codes for small ribosomal subunit</p></li><li><p>cloesly related species (same genus) have similar rRNA sequences </p><ul><li><p>info can be used to make phylogenetic trees using computer-based software</p></li></ul></li><li><p>can also be used as qualititative method to determine unknown species and diff types in sample </p></li></ul><p></p>
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advantages of sequencing rRNA

  • all living organisms need ribosomes to make proteins hence rRNA gene is in all species

  • rRNA gene has highly conserved regions where few mutations occur - researchers can synthesise primers that will bind in complementary fashion so PCR can be done

  • also consists of multiple highly variable regions where mutations occur more readily - researchers can distinguish between diff species

  • can detect many species at once in comparison to DNA sequencing of individual species - useful since theres >billion prokaryote species

  • DNA sequencing requires prokaryote species to be isolated and some cant be cultured and hence isolated easily


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protein sequencng'/aa seuquncint


  • if diff species produce proteins w/ v similar aa sequences, DNA is v similar, inherited from recent common ancestor

  • proteins maintained through evolutionary history are identified (those crucial to survival)

    • e.g. cytochrome c - needed for aerobic respiration

    • protein varies from species to another - degree of similarity indicates closeness of evolutionary relationship

    • assumed that 1st organism had cytrochrome c, over billions of yrs mutations changed sequence of bases in gene coding for protein


<ul><li><p>if diff species produce proteins w/ v similar aa sequences, DNA is v similar, inherited from recent common ancestor</p></li><li><p>proteins maintained through evolutionary history are identified (those crucial to survival)</p><ul><li><p>e.g. cytochrome c - needed for aerobic respiration</p></li><li><p>protein varies from species to another - degree of similarity indicates closeness of evolutionary relationship</p></li><li><p>assumed that 1st organism had cytrochrome c, over billions of yrs mutations changed sequence of bases in gene coding for protein</p></li></ul></li></ul><p></p>
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DNA-DNA hybridisation process

  1. particular gene sequence to be compared between species is isolated using gene probe and enzymes

  2. DNA of 2 species is heated to 95 to seperate strands

  3. strands are mixed togehter and allowed to cool - enables formation of hydrid DNA

  4. strands are reheated to work out ‘melting pt’ of hybrid DNA. melting pt is defined as temp where 50% of DNA is single stranded, other half is double. higher melting pt, more bonding between DNA molecules, more in common 2 DNA sequences are

  5. higher the melting point, the greater similarity between the DNA sequences and the more likely it is that they shared a more recent common ancestor


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DNA DNA hybridisation general info

  • heating solution containing DNA disrupts H bonding between 2 complementary strands (measured as boiling pt)

  • when mixture is cooled, they recombine to form double helix again

  • solutoin of DNA is heated from diff species is heated, allows hybrid DNA to form

  • degree of H bonding between one species DNA and another gives measure of similarity of sequence of nucleotides, indicator of how closely related species are


<ul><li><p>heating solution containing DNA disrupts H bonding between 2 complementary strands (measured as boiling pt)</p></li><li><p>when mixture is cooled, they recombine to form double helix again</p></li><li><p>solutoin of DNA is heated from diff species is heated, allows hybrid DNA to form </p></li><li><p>degree of H bonding between one species DNA and another gives measure of similarity of sequence of nucleotides, indicator of how closely related species are </p></li></ul><p></p>
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principal groups for classifying organisms:

domain

kingdom

phylum

class

order

family

genus

species


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naming species

  • binomial nomenclature - first name of species is genus, second is species name

  • genus name is capitalised

  • both written in italics on typing, if handwritten underlined


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biochemical similarities

  • DNA is universal

  • biochemical similarities of DNA and proteins can be used to determine between asequally reproducing species

  • also other biochemical molecules that are v similar between soecies

    • cellular respiration, photosynthesis, transcription, translation are essential processes in organisms ∴ biomolecules involved in processes are similar

    • e.g. cytochrome c (cellular respiration)

    • similarity can be compared to determine evolutaionry relationships


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population definition

group of individuals of the same species occupying the same space and interbreeding

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gene pool

  • sum of all genes (alleles) of all individuals in a population - form which guture generations of individuals will be created

  • populations with large gene pools (large diersity of alleles) will be more robust, more likely to survive changing or harsh environmental conditions

  • reduced genetic diversity means greater extinction risk


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biological species concept

  • most widely used, based on reproductive isolation

  • members of one species are a population where members have similar characteristics (biochemical, morphological, and genetic) and capacity to interbreed and produce fertile offspring

  • species consist of several populations that are geographically isolated

  • even though physically isolated, still considered one species if contribute to gene pool by producing fertile offspring


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limitations of biological species concept - asexually reproducing species

  • doesnt apply to them (e.g. bacteria, hydra, sea stars)

  • biochemical comparisons of DNA and proteins help establish relatedness in this circumstance


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limitations of biological species concept - lack of data on breeding behaviour

  • many speciies dont have extensive research conducted on breeding patterns and ability to produce fertile offspring w/other groups of organisms


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limitations of biological species concept - fossils

  • DNA of fossils has generally degraded for species that are long extinct - rules out ability to compare breeding behaviours and comparative genomics.

  • also no evidence regarding interbreeding and behaviour of fossilised organisms.

  • can still show structural (morphological) featers, which can be compared



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limitations of biological species concept - overlap of populations and hybrids

  • difficult yo examine extent of interbreeding

  • overlap in distribution areas and presence of hybrids exists

  • if diff give rise to intermediate forms, considered one species

  • may classify into seperate subspecies


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morphological species concept

  • groups species according to structural features that are unique to particualr group of organisms

  • can be applied to any group, mainly useful for fossils


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limiation of morphological species concept

  • different species have similar structural features

  • thus, only used when biological species concept cant be used

  • species w/ similar features due to convergent evolution is example of the limit


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fossils

  • DNA in fossils generally degraded for species that are extinct for long time - cant compare breeding or comparative genomics, no evidence of interbreeding

  • can still show structural features - thus morphological species concept used


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reproductive isolation

  • members of separate species cannot interbreed and have fertile offspring

  • achieved through pre and post zygotic mechanisms

  • pre - occur before zygote formed (before fertilisation)

  • post - occur after zygote formed, prevent from developing into viable, fertile adult hybrid


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pre-zygotic mechanism - temporal (time) isolation

  • breeding cycles/time of fertility differs between one population of species to the next

  • e.g. plants flower at different seasons, some animals are nocturnal and others arent


<ul><li><p>breeding cycles/time of fertility differs between one population of species to the next</p></li><li><p>e.g. plants flower at different seasons, some animals are nocturnal and others arent</p></li></ul><p></p>
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pre-zygotic mechanism - behavioural isolation

  • development in populations of highly specific calls or rituals detected and responded only by other members of particular populatoin

  • e.g. mating calls, mating dances


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pre-zygotic mechanism - mechanical (morphological) isolation

  • incompatible sexual organs, whereby significant anatomical differences act as powerful mecahnism to ensure different populations remain reproductively isolated

  • eg. of plants - use different populations


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pre-zygotic mechanism - gamete isolation

  • incompatibility of sperm and eggs from populations of different species

  • range of mechanisms and chemicals in species ensuring sperm can fertilise and fuse with egg from member of same species only

  • isolating mechanisms operating here include:

    • sperm does not respond to chemical signals from egg

    • sperm cannot penetrate egg


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post-zygotic mechanisms - hybrid inviability

  • hybrid is weak and compete poorly for resoruces

  • unlikely to reach reproductive age and contribute to gene pool


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post-zygotic mechanisms - hybrid sterility

  • extension of hybrid inviability

  • hybrid formed is sterise, hence cannot contribute to gene pool

  • chromosome number of 2 parents differs, hence do not have complete set of homologous pairs and meitoci division cannot occur in normal fashion


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dominant and recessive, hetero and homozygous explained

  • zygote inherits ½ genetic material from each parent

  • genes act in various ways - some according to dominant and recessive patern of inheritence

  • homozygous (2 dominant or 2 recessible

  • heterozygous (1 dominant 1 recessive)

  • dominant - alleles that are expressed when at least one dominant gene is in genotype

  • recessive - 2 copies required to be expressed


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cystic fibrosis and type of mutation

  • caused by faulty recessive allele of a gene

  • normal (C) is dominant over recessive mutant allele ©

  • CC - normal, does not carry CF allele

  • Cc - normal, carries one CF ellele

  • cc - CF sufferer, has 2 CF alleles

  • lethal mutation


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sickle-cell anaemia caused how and homo and heterozygous

  • change in one base, leads to protein with one aa changed

  • A is changed to T in substitution mutation in codon for 2th aa in beta chain; codon changes from GAG to GTG

    • causes glutamic acid to be replaced by valine

  • normal haemoglobin HbA and mutatn HbS alelles therefore:

    • homozygous can be HbA HbA or HbS HbS

    • heterozygous is HbA HbS


<ul><li><p>change in one base, leads to protein with one aa changed</p></li><li><p>A is changed to T in substitution mutation in codon for 2th aa in beta chain; codon changes from GAG to GTG</p><ul><li><p>causes glutamic acid to be replaced by valine</p></li></ul></li><li><p>normal haemoglobin HbA and mutatn HbS alelles therefore:</p><ul><li><p>homozygous can be HbA HbA or HbS HbS</p></li><li><p>heterozygous is HbA HbS</p></li></ul></li></ul><p></p>
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sickle cell anaemia sufferers how does it work (and codominance)

  • HbA and HbS exhibit form of co-dominance (or incomplete dominance) - heterzygous form exhibits features from both alleles (normal and abnormal haemoglobin made)

  • people with mutant gene and mutant heamoglobin dont have normal bi-concave TBC - become sickle-shaped

    • often causes death in homoxygous individiuals


<ul><li><p>HbA and HbS exhibit form of co-dominance (or incomplete dominance) - heterzygous form exhibits features from both alleles (normal and abnormal haemoglobin made)</p></li><li><p>people with mutant gene and mutant heamoglobin dont have normal bi-concave TBC - become sickle-shaped</p><ul><li><p>often causes death in homoxygous individiuals </p></li></ul></li></ul><p></p>
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types of mutations:

  • lethal

  • disadvantageous

  • neutral

  • beneficial


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disadvantageous mutation e.g. osteoporosis

  • cause disease in those with mutant gene but is not fatal

  • e.g. osteoporosis

    • sufferers have brittle bones

    • low-density lipoprotein receptor related protein 5 (LRP5) involved in bone density and mutations in gene can cause osteoporosis

    • bones of affected individuals lose/’leak’ minerals (e.g. calcium) into blood


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neutral mutations

  • no affect on either appearence (phenotype) or its function

  • majority of mutations are neutral

    • some scientists argue they are still mutations - change genotype and will be passed to guture generations (if germline cells)

    • changes to introns of DNA, may cause unknown changes to function


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sickle cell anaemia

  • homozygous individuals have LETHAL mutation

  • heterozygous individuals (HbA, HbS) have some HbS-derived haemoglobin but more normal haemoglobin - suffer form mild anaebia

  • if in malaria infested area, protists that cause disease cannot reproduce in RBC with haemoglobin made from HbS allele

    • have reduced chance of getting malaria


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sexual reproduction on evolution (and evolution in asexually reproducing)

  • introduces variety into offspring so they dont look the same as parents

  • variation expresses itself as diff genotypes (pairs of alleles of genes) and thus different phenotypes (appearance)

  • mutations are only source of new alleles in genes, sexual reproduction reshuffles alleles of genes in parents to give new combos

  • popultions of sexually reproducing species show variation between individual members of population

  • asexual - reproduction happens quickly if there is a favourable mutation


<ul><li><p>introduces variety into offspring so they dont look the same as parents</p></li><li><p>variation expresses itself as diff genotypes (pairs of alleles of genes) and thus different phenotypes (appearance)</p></li><li><p>mutations are only source of new alleles in genes, sexual reproduction reshuffles alleles of genes in parents to give new combos</p></li><li><p>popultions of sexually reproducing species show variation between individual members of population</p></li><li><p>asexual - reproduction happens quickly if there is a favourable mutation</p></li></ul><p></p>
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how variation introduced in meiosis

  • crossing over - introduces new combo of alleles of maternal and paternal genes

  • independent assortment of chromosomes - gives rise to new combos of maternal and paternal chromosomes in gamets through random alignment of homologous chromosomes on opposite side of equator of cell (meiosis 1)


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how variation introduced in fertilisation

  • random fusion of an ovum and sperm, each w/ unique collection of chromosomes and alleles


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allele frequency

  • relative proportion of particular allele in population

  • usually expressed as fraction or percentage


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changes in gene pool bought on by:

  • mutations

  • gene flow

  • genetic drift


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changes in gene pool - gene flow

  • movement of alleles between diff populations of same species

  • occur when individuals move in and out of populations, or migrate - change frequency of alleles found in gene pool

  • inc. flow of alleles of genes between populations reduces diff between populations

  • limited gene flow leads to populations evolving and developing in own environment


<ul><li><p>movement of alleles between diff populations of same species </p></li><li><p>occur when individuals move in and out of populations, or migrate -  change frequency of alleles found in gene pool</p></li><li><p>inc. flow of alleles of genes between populations reduces diff between populations</p></li><li><p>limited gene flow leads to populations evolving and developing in own environment</p></li></ul><p></p>
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changes in gene pool - genetic drift

  • random changes in frequency of alleles, more pronounced effects on small populations

  • if any individuals dont contribute their alleles to next gen, major impact of gene pool

  • seen in populations where effects (e.g. bottlneck) occur

  • reduced population size reduces variation in gene pool → reduced genetic biodiversity


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bottleneck affect

  • impact of random factors (e.g. bushfires) on population

  • changes allele frequency - removes certain alleles due to random cahnce


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natural selection and darwins hypothesis

  • for each population, more individuals were born and survived to reproduce, and environmental factors hold all populations in check

  • natural selection - nature/the environment selected individuals who were best adapted -

  • variation between individuals gave reproductive advantages, enable organisms to have more offfspring to survive next gen

  • life evolved from common acnester, and due to natural slection, populations were modifed over time according to diff environments and diff selective pressures they were exposed to


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darwins finches

  • have diff beak shapes - provide unique adaptation for type of food predominantly found in particular environment of diff islands


<ul><li><p>have diff beak shapes - provide unique adaptation for type of food predominantly found in particular environment of diff islands</p></li></ul><p></p>
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natural selection def and selective pressures

  • natural selection - nature/the environment selected individuals who were best adapted

  • selective pressures act on phenotypes of individuals in the population, leading to changes in in allele frequency

    • certain alleles are favoured, otehrs are selected against

  • ‘survival of the fittest’ - alleles of favoured traits in population of gene pool inc in frequency over time

  • diff selection pressures can lead to divergent evolution or adaptive radiation


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biotic selection factors

  • predators

  • disease-causing organisms

  • competition from members of the same or different species

  • symbiotic relationships with other organisms

  • human activities


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abiotic selection factors

  • rainfall

  • temperature

  • nutrient levels

  • light intensity

  • medical drugs (e.g. antibiotics)

  • chemicals (e.g. pesticides)


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industrial melanism in peppered moths

  • before industrial revolution, trees moths lived in were covered in offwhite lichen

  • moths were light (light work) and speckled - camouflaged from predation by birds

    • occasionally black (dark form) would grow - high visibility, high chance of being eaten before reproducing

  • due to coal based industry, trees covered in black soot - white moths were easily spotten and eaten

    • black (melanic) variant became predominant by 1850 - phenomenon called industrial melanism

  • reduction of coal use in later 20th century made trees green (algae) - now both moth forms are common - balanced polymorphism


<ul><li><p>before industrial revolution, trees moths lived in were covered in offwhite lichen </p></li><li><p>moths were light (light work) and speckled - camouflaged from predation by birds </p><ul><li><p>occasionally black (dark form) would grow - high visibility, high chance of being eaten before reproducing</p></li></ul></li><li><p>due to coal based industry, trees covered in black soot - white moths were easily spotten and eaten</p><ul><li><p>black (melanic) variant became predominant by 1850 - phenomenon called<strong> industrial melanism </strong></p></li></ul></li><li><p>reduction of coal use in later 20th century made trees green (algae) - now both moth forms are common - <strong>balanced polymorphism</strong></p></li></ul><p></p>
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summary of evolution by natural selection

  • individuals in populations show phenotypic variation - result of environment and differences in genotype (inherited by next gen)

  • struggle for suvial between individuals of population due to biotic or abiotic selection pressures

  • organisms best adapted to environmental conditions are more likely to survive to reproductive age and pass favourable alleles of genes to next gen

  • over many gens, populations may evolve (showing changes in gene pool)

    • effects of natural selection can be measured by changes in frequency of alleles in gene pool - evidence of evolution occuring

    • populations/species that evolve, not individual organismsm

  • environmental elective pressures determine reproductive success of population, population does not adapt to conditions


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process of evolutino suggests

  • all life forms evolved from single, simple common ancestor (LUCA)

  • species change through time

  • evolution in a species can be recognised by changing frequency of alleles in population


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founder effect

  • small group of individuals separate/move away from original population

  • founding populations have a very different genotype from the original

  • changes allele frequency in populations


<ul><li><p>small group of individuals separate/move away from original population</p></li><li><p>founding populations have a very different genotype from the original </p></li><li><p>changes allele frequency in populations </p></li></ul><p></p>
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speciation

  • process that gives rise to new species

    • gene flow between 2 populations has stopped and reproductive isolation occurs

    • causes change in allele frequency in each population at differen rates

    • cumulative change in allele frequency over time means populations cant meet biological species def

  • 2 types: allopatric and sympatric


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allopatric speciation

  • Members of the original population are separated by a geographical barrier - gene flow between them is prevented

  • each population is subjected to different biotic and abiotic factors - selective factors and natural selection acts diff in each new location

  • genetic diff accumulate in isolated populations

  • if gene pools are separated and no gene flow occurs for long enough, geographical isolation evolves to reproductive isolation

    • speciation has occurred by divergent evolution

  • adaptive radiation can occur


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adaptive radiation

  • A single ancestral species rapidly diversifies into new forms to fill different ecological niches

  • divergent evolutoin of related species from common ancestor



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sympatric speciation

  • one population gives rise to two or more new species while inhabiting the same region (no geographical isolation)

  • gene flow must be prevented/reduced - reproductive isolatino must ultimately arise

  • occurs via:

    • occupying different microhabitats

    • polyploidy


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sympatric speciation - occupying different microhabitats

  • particular members of one population occupy specific niche or microhabitat

  • creates ecological isolation within same geographical area

    • diff microhabitats impose contrasting environmental pressures - favour traits for each speficic niche

    • habitat based mating

    • causes gene flow restriction into the subpopulations


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sympatric speciation - polyploidy

  • organisms possess more than two sets of chromosomes

  • occurs commonly in plants and certain groups of fish and amphibians

  • result of malfunction during meiosis - chromosomes fail to separate properly (nondisjunction)

  • example of instantaneous speciation - polyploid organism can’t interbreed with organism with a normal diploid number; gene flow is stopped

  • major role in flowering plant evolution


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convergent evolution

  • different species of roganisms develop similar (analogous) features with similar functions

  • different groups of organisms (often in similar habitats w/similar selective pressure) have v similar structures/behavoiurs

  • occurs when analogous features evolve independently in unrelated groups or seperate species


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analogous features

  • evolved separately in different lines of evolution

  • similar functoins, different anatomy

  • formed by convergent evolution


<ul><li><p>evolved separately in different lines of evolution</p></li></ul><ul><li><p>similar functoins, different anatomy </p></li><li><p>formed by convergent evolution</p></li></ul><p></p>
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convergent evolution - wings and eyes

wings:

  • birds, bats, and butterflies (unrelated) all have wings for flight - invovled independently of each other


eyes:

  • an ancestral master gene ‘Pax6’ - og gene responsible for development of collections of simple light-sensitive cells

  • found across diverse set of organimss

  • probably part of genome of very early comon ancestor lived approx 500mil yrs ago

  • cephalopods and vertebrate have camera eye


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divergent evolution

  • evolution of species from common ancestor

  • groups separate by speciation - accumulation of diff in geome by mutation

  • subjected to diff selective pressures therefore no gene flow - reproductively isolated

  • have homologous features


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homologous features

  • share a common ancestor, have physical features with common evolution that have different functions

  • e.g., pentadactyl limb


<ul><li><p>share a common ancestor, have physical features with common evolution that have different functions</p></li><li><p>e.g., pentadactyl limb </p></li></ul><p></p>
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divergent vs convergent evolution comparison

knowt flashcard image
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succession

  • gradual process where the species composition of a community changes and is brought about by the modification of the habitat caused by organisms that live there

  • leads to significant change that whole ecosystem changes to favour new mix of species - caused by long term change due to colonisation or natural disasters


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ways organisms modify ecosystems

  • provide food or shelter for others

  • changing soil structure, causing breakdown of rock or increasing organic matter in soil

  • decomposing dead organic material, making nutrients available to other organisms


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primary succession

  • unidirectional process - lifeless ecosystems are colonised by progressively more stable communities of living things

  • 1st stage - colonisation of bare land by pioneer species

    • carry out life processes (inc chemical reactions weathering rock over time)

    • transform material to soil

    • death and decay, increasing nutrient content in soil

  • intermediate species sprout from seeds blown from nearby areas or insects and birds

  • area is colonised by plants that become communities dominant vegetation - climax community produce


<ul><li><p>unidirectional process - lifeless ecosystems are colonised by progressively more stable communities of living things</p></li><li><p>1st stage - colonisation of bare land by pioneer species</p><ul><li><p>carry out life processes (inc chemical reactions weathering rock over time)</p></li><li><p>transform material to soil</p></li><li><p>death and decay, increasing nutrient content in soil</p></li></ul></li><li><p>intermediate species sprout from seeds blown from nearby areas or insects and birds</p></li><li><p>area is colonised by plants that become communities dominant vegetation - climax community produce</p></li></ul><p></p>
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adaptations that improve survival of pioneer species

  • autotrophic nutrition

  • ability to reproduce quickly

  • ability to tolerate harsh environmetal conditions


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secondary succession

  • occur in mature ecosystems following natural disaster or major human impact (e.g. forest clearing)

  • e.g. woodland ecosystems destroyed by wildfires

    • fire stimulates germination of seeds; growth is more rapid than with primary succession - soils are warm, high nutrient content, can retain water better

    • new mix of plants and animal populations is quickly established

    • competition between colonising species populations and returning and new species influences new community and how it compares to og

  • different selective pressures can lead to divergent evolutoin or adaptive radiation as species exploit new niches


<ul><li><p>occur in mature ecosystems following natural disaster or major human impact (e.g. forest clearing)</p></li><li><p>e.g. woodland ecosystems destroyed by wildfires</p><ul><li><p>fire stimulates germination of seeds; growth is more rapid than with primary succession - soils are warm, high nutrient content, can retain water better</p></li><li><p>new mix of plants and animal populations is quickly established</p></li><li><p>competition between colonising species populations and returning and new species influences new community and how it compares to og</p></li></ul></li><li><p>different selective pressures can lead to divergent evolutoin or adaptive radiation as species exploit new niches</p></li></ul><p></p>
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characterisits of mature or late sucession in communities and ecosystems

  • usually more species

  • number of heterotrophs increases more than number of autotrophs

  • recycling of nutrients is more efficient

  • organisms are more specialised


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importance of species diveristy

  • species w/ reduced gene pool have higher risk of extinction - greater risk of becoming extinct due to greater susceptibility to environmental pressures


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affect of human population growht on biodiversity

  • exponential growth of global human population

  • unsustainable - huge demands on resources needed by humans - jeopardises biodiversity


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introduced/invasive/exotic species

  • foreign species brought to australia from other countries - move intentionally or accidentallyform native location to new geographic area

  • those who cause harm are invasive

  • compete with native species for habitat or resources or directly feed on them

  • if they have virtually no natural predators, spread across huge areas and reduce biodiversity


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introduced species eg. - cane toads and rabbits

cane toads:

  • native to Central and south america

  • introduced to QLD to act is biological control agents of cane beetles

  • have virtually no natural predators - spread over huge area of Australia

  • Predators die after ingesting the toad due to poison glands on their backs - reduce biodiversity


rabbits:

  • destroy native vegetation, compete with native animals for resources

  • take over burrows from burrowing Australian species (eg. bandicoots, bilby)


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serious threats of pollution

  • chemical pesticides affect higher order consumers through biomagnification (DDT, insecticides)

  • oil spills from ships at sea

  • accumulation of plastic waste in waterways and oceans

    • kills animals

    • degradation into microplastics - enter food chain through biomagnification

  • acid rain and photochemical smog - product of emission of SO2 and NOx into atmosphere from fossil fuel combustion

  • ineffective or insufficient sewage treating and disposal

  • production, transport, and storage of radioactive waste

  • impact of chlorofluorocarbons (CFCs) used in aerosols and fridges - destroys ozone layer