Crude birth rate (CBR)
(total births in a year / total population) x 1000
Crude death rate (CDR)
(total deaths in a year / total population) x 1000
Natural increase rate (NIR)
(CBR - CDR) / 10
Doubling time (DT)
70 / NIR (%)
Ecological footprint (EF)
total area required to sustain a population / total population size
Net primary productivity (NPP)
NPP = GPP - R
Gross secondary productivity (GSP)
Food eaten - fecal loss
Net secondary productivity (NSP)
GSP - R
Trophic level efficiency
(energy at higher trophic level / energy at lower trophic level) x 100
Biochemical oxygen demand (BOD)
Amount of dissolved oxygen needed by microorganisms to decompose organic matter (higher indicates more pollution)