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Microevolution
The change in allele frequencies within a population over generations.
Speciation
The evolutionary process by which one species splits into two or more species.
Macroevolution
Large-scale evolutionary change above the species level, affecting major taxonomic groups.
Radiometric dating
A method for determining the age of objects using the known, steady decay rates of radioactive isotopes.
Radiocarbon dating
A radiometric dating method using carbon-14 with a 5730-year half-life; used for organic remains up to ~60,000 years old.
Electron spin resonance dating
“Trapped-electron” dating used to determine ages of materials like tooth enamel; effective up to 5 million years.
Potassium-Argon dating
A radiometric technique using the decay of K-40 to Ar-40 to date volcanic rock up to billions of years old.
Stromatolites
Living mounds of mineralized cyanobacterial biofilms; the oldest known fossils (~3.5 billion years old).
Protocell
A membrane-bound collection of organic molecules thought to be the precursor to true cells.
Ribozymes
RNA molecules capable of catalyzing chemical reactions, including self-replication.
Miller-Urey experiment
An experiment simulating early Earth conditions that produced organic molecules, including amino acids.
Plate tectonics
The theory that Earth’s crust is divided into plates that float and move due to convection in the mantle.
Continental drift
The gradual movement of Earth's continents over geological time.
Mass extinction
A rapid and widespread loss of biodiversity marking transitions between major geological eras.
Adaptive radiation
Rapid diversification in which a single ancestor gives rise to many new species, often after ecological opportunities arise.
Biological species concept
Defines species as groups whose members interbreed in nature and produce fertile offspring; relies on reproductive isolation.
Reproductive isolation
Physical, biological, or behavioral barriers preventing species from interbreeding.
Allopatric speciation
Speciation that occurs when populations are geographically separated.
Sympatric speciation
Speciation that occurs in populations living in the same area due to ecological or genetic factors.
Polyploidy
Condition of having more than two sets of chromosomes; often causes instantaneous plant speciation.
Hybrid
Offspring resulting from the mating of individuals from two different species.
Hybrid vigor
When hybrids show superior traits (e.g., increased fitness) compared to parent species.
Habitat isolation
A pre-zygotic barrier where species occupy different habitats and rarely encounter one another.
Temporal isolation
A pre-zygotic barrier where species breed at different times (season, day, or year).
Behavioral isolation
A pre-zygotic barrier where differences in courtship behaviors prevent mating.
Mechanical isolation
A pre-zygotic barrier where morphological differences prevent mating or fertilization.
Gametic isolation
A pre-zygotic barrier where gametes (egg & sperm) cannot fuse due to molecular incompatibility.
Reduced hybrid viability
A post-zygotic barrier where hybrids fail to develop properly or die early.
Reduced hybrid fertility
A post-zygotic barrier where hybrids survive but are sterile (e.g., mule).
Hybrid breakdown
A post-zygotic barrier where first-generation hybrids are fertile but their offspring are weak or sterile.
Fossil record
The chronological collection of life’s remains showing historical changes in species over time.
Biogeography
The study of the geographic distribution of species.
Homology
Similarity in structure due to shared ancestry.
Homologous structures
Structures with similar internal anatomy but different functions due to shared ancestry (divergent evolution).
Analogous structures
Structures with similar function but different ancestry; result of convergent evolution.
Convergent evolution
Independent evolution of similar traits in unrelated organisms due to similar environmental pressures.
Artificial selection
Selective breeding by humans to increase desirable traits in organisms.
Population
A group of individuals of the same species living in a specific geographic area.
Gene pool
All alleles present in a population.
Genetic variation
Differences in DNA sequences between individuals of a population.
Hardy-Weinberg equilibrium
A state where allele frequencies remain constant if no mutation, migration, drift, non-random mating, or selection occurs.
Mutation
A change in DNA sequence; the ultimate source of new genetic variation.
Genetic drift
Random changes in allele frequency, strongest in small populations.
Founder effect
When a few individuals start a new population with allele frequencies different from the original.
Bottleneck effect
When a population undergoes a drastic reduction in size, altering allele frequencies.
Gene flow
Movement of individuals (and their alleles) between populations.
Natural selection
Process where better-adapted individuals survive and reproduce, increasing advantageous alleles.
Evolutionary fitness
An individual’s ability to contribute viable, fertile offspring to the next generation.
Relative fitness
Fitness measured relative to others in the population.
Sexual selection
A type of natural selection based on traits that increase mating success.
Sexual dimorphism
Differences in appearance between males and females of the same species.
Intrasexual selection
Competition between individuals of the same sex (usually males) for mates.
Intersexual selection
Mate choice, often where females select males based on traits or displays.
Directional selection
Selection that favors one extreme phenotype.
Disruptive selection
Selection that favors both phenotypic extremes and acts against intermediates.
Stabilizing selection
Selection that favors intermediate phenotypes and reduces extremes.
Theory of evolution
The concept that species change over time and modern species descend from ancestral ones.
Adaptation
A heritable trait that increases survival and/or reproduction in a specific environment.
Phylogeny
Evolutionary relationships among organisms
Taxonomy
Biological classification
Shared derived character
Distinguishing feature of a group that is not shared by previous ancestors
Claude
Group of organisms composed of common ancestor and all descendants
Vertebral column
Support structure, also protects nerve cord
Vertebrates
Organisms with a skull and vertebral column
Chondrichthyans
Cartilaginous fish; sharks and rays with derived character of jaws and paired appendages
Ray-finned fish
Greatest number of species of any vertebrate; derived character of internal bony skeleton, lungs/lung derivative
Lobe-finned fish
Bony; derived character of loved appendages (precursor to limbs)
Tetrapod
“Four feet;” jawed vertebrates that have limbs and feet that can support their weight on land
Convergent evolution
Unrelated species develop similar traits due to similar environmental pressures
Divergent evolution
Closely related species develop different traits as they adopt to different environments
Analogous structures
Similar function, different origin
Homologous structures
Shared origin, different function
Amphibians
Organisms that have legs with digits; first vertebrates to colonize land
The Sixth Mass Extinction
Climate change, deforestation, overharvesting
Reptiles
Organisms with an amniotic egg; capable of completing life cycles without “attachment” to water
Ectothermic
Absorb external heat rather than generating most of their own
Mammals
Organisms with hair and mammary glands producing milk
Monotremes
Oldest lineage of mammals; lay eggs
Marsupials
Nurse in external pouch after short pregnancy
Placental mammals
Placenta provides nutrients from mother
Apes
Species with larger brains relative to body size compared to monkeys and other primates
Hominins
Group including modern humans and extinct human ancestors
Homo erectus
First hominins to migrate out of Africa; first humans to use fire
Homo neanderthalensis
Last common ancestor with humans 400,000 years ago. Interbred with Homo sapiens: non-Africans roughly 2% Neanderthal DNA
Homo sapiens
Originated in Africa, migrated all over the world. Highest levels of brain cognition and adaptability; sophisticated use of tools and fire.