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The closest protist relatives to animals
Both are unicellular or colonial with a collar of microvilli around a flagellum.
Similar cell signaling and adhesion genes.
Morphological resemblance to choanocytes (collar cells) in sponges.
choanoflagellate
Main Stages of Animal Embryonic Development
Zygote: Fertilized egg.
Cleavage: Rapid mitotic divisions without growth.
Blastula: Hollow ball of cells with a fluid-filled cavity (blastocoel).
Gastrulation: Cells migrate inward to form germ layers and the archenteron (primitive gut).
Gastrula: Embryo with ectoderm, mesoderm (in triploblasts), and endoderm.
Hox genes: A family of regulatory genes that control body plan development along the anterior-posterior axis.
Earliest Animal Fossils
Ediacaran biota.
Dated to about 558 million years ago
Radial symmetry
Symmetry around a central axis (e.g., jellyfish)
Suited for sessile or drifting lifestyles, can interact with environment from all sides.
Bilateral symmetry
Left and right halves mirror each other (e.g., humans).
Cephalization (head), directional movement, suited for active lifestyles
Asymmetry
No symmetry (sponges)
Triploblastic Body Cavity Types
Coelomates
Acoelomates
Pseudocoelomate
Coelom:
Fluid-filled body cavity lined by mesoderm.
Coelomates:
Animals with a true coelom (e.g., annelids).
Acoelomates:
No body cavity (e.g., flatworms).
Pseudocoelomates:
Body cavity not fully lined by mesoderm (e.g., roundworms).
Hemocoel:
Primary body cavity in some invertebrates (e.g., arthropods), filled with hemolymph.
Hemolymph
Circulatory fluid in organisms with open circulatory systems.
Protostomes
Cells are spiral and determinate—each cell’s fate is fixed early
blastophore becomes mouth
Deuterostomes
Cells are radial and indeterminate—each cell can develop into a complete embryo if separated
blastophore becomes anus
Phylum Porifera
sponges
Eumetazoa
Includes all animals with true tissues, excludes sponges
How do sponges feed
filter feeding
water enters through pores, flows into the spongocoel, and exits via the osculum. Choanocytes trap food particles with their flagella
Epithelial cells (pinacocytes)
outer covering, regulate surface area of sponge
Pores (ostia)
water enters sponge
Spongocoel
central cavity of sponge
Osculum
large opening for water exit sponge
Choanocytes
collar cells, trap food
Amoebocytes
attached to collar cells, digest food, transport nutrients, form spicules
Mesohyl
gelatinous matrix between cell layers in sponge
Spicules
structural support, deter predators of sponge
Sequential hermaphroditism
an individual functions as male and then female (or vice versa) at different times. This increases reproductive success when mates are scarce
How sponges help people
Sponges produce bioactive compounds used in cancer, HIV, and antibacterial drugs
Cnidaria
jellyfish, corals, sea anemones.
Cnidarian body forms
Polyp: sessile, cylindrical (e.g., sea anemone).
Medusa: free-swimming, bell-shaped (e.g., jellyfish).
They have one opening to the gastrovascular cavity
Cnidocytes
stinging cells
nematocysts
inject toxins to capture prey or defend
Do cnidarians have a brain
No brain. They have a nerve net—a decentralized web of neurons that allows basic responses to stimuli
Bilateria
Deuterostomia, Ecdysozoa, Lophotrochozoa
Lophotrochozoa
Lophophore: ciliated feeding structure.
Trochophore: larval stage with cilia.
Six Lophotrochozoa
Platyhelminthes (e.g., planaria)
Rotifera (e.g., rotifers)
Nemertea (e.g., ribbon worms)
Mollusca (e.g., snails, molluscs)
Annelida (e.g., earthworms)
Brachiopoda (e.g., clams).
Planaria (flatworm)
belong to Platyhelminthes. They can regenerate entire bodies from small fragments using pluripotent stem cells
have a centralized nervous system with ganglia and nerve cords, unlike cnidarians’ nerve net
Tapeworms
in Platyhelminthes. They absorb nutrients directly through their skin from the host’s intestine
Parthenogenesis
reproduction without fertilization. In rotifers, females can reproduce asexually, allowing rapid population growth
Lophophore:
a crown of ciliated tentacles used for filter feeding in Ectoprocta and Brachiopoda
Mollusk body
Foot (movement)
Visceral mass (organs)
Mantle (secretes shell)
Radula
a chitinous, toothed ribbon used to scrape or cut food (not found in bivalves)
mouth in molluscs
Mollusca
snails, clams, squids, octopuses
Only cephalopods (e.g., octopus) have a closed circulatory system, which is more efficient for active movement
Mollusk extinction causes
habitat loss, pollution, climate change, invasive species, and overharvesting
Cnidarians vs Molluscs
Cnidarians have a gastrovascular cavity with one opening. Mollusks have a complete digestive tract with separate mouth and anus
Ecological and Economic Importance of Fungi
• Decomposers: Fungi break down dead organic matter, recycling nutrients like carbon and nitrogen.
• Pathogens: Some fungi cause diseases in plants (e.g., rusts, smuts) and humans (e.g., Candida, Aspergillus).
• Mutualists:
• Mycorrhizae: Symbiosis with plant roots enhances nutrient uptake.
• Lichens: Symbiosis with algae/cyanobacteria, colonizing harsh environments.
• Beneficial Uses: Antibiotics (penicillin), food (yeast, mushrooms), bioremediation, and industrial enzymes.
Unicellular Fungi
Yeasts
Chytridiomycota
Aquatic, flagellated spores
Batrachochytrium
Zygomycota
Coenocytic hyphae, zygospores
Rhizopus (bread mold)
Glomeromycota
Arbuscular mycorrhizae
Symbionts with plants
Ascomycota
Asci with ascospores
Saccharomyces (yeast)
Basidiomycota
Basidia with basidiospores
Agaricus (mushroom)
Mycorrhizae
Found in soil, enhance plant nutrient uptake
Lichens
Found on rocks, trees, soil; bioindicators of air quality
Asexual Reproduction in Fungi
Budding (yeast), fragmentation, spore formation (conidia, sporangiospores).
Mating Types vs. Sexes
Fungi use mating types (e.g., +/−) instead of male/female. This reflects genetic compatibility, not gamete size or sex roles
Fungal Sexual Reproduction
Plasmogamy → Dikaryotic stage → Karyogamy → Diploid zygote → Meiosis.
Asexual spores
Mitosis, genetically identical, rapid spread
Sexual spores
Meiosis, genetically diverse, formed after plasmogamy/karyogamy
Fungal Life Cycle Stages
Haploid (n): Most of life cycle.
Heterokaryotic (n + n): Two nuclei coexist.
Diploid (2n): Brief, after karyogamy.
Heterokaryotic stage is unique to fungi
Lichen Forms
Crustose: Flat, crusty.
Foliose: Leaf-like.
Fruticose: Shrubby, branched
Soredia and Lichen Reproduction
Soredia: Asexual propagules (fungal hyphae + algal cells).
Dispersed to form new lichens.
Asexual reproduction