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Apical Meristem
Region of actively dividing cells responsible for primary growth
Lateral meristem
Cylindrical meristem responsible for secondary growth
Shoot apical meristem (SAM)
Produces stems, leaves axillary buds, floral structures
Root apical mersitem (RAM)
Produces the primary tissues of the root
Primary growth
Increase in root or shoot length
Secondary growth
Increase in stem or root diameter
Dermal tissue system
Epidermis and related tissues protecting the plant surface
Ground tissue system
Parenchyma, collenchyma, and sclerenchyma tissues
Vascular tissue system
Xylem and phloem responsible for long distance transport
Epidermis
Outermost protective cell layer of primary plant organs
Cuticle
Waxy surface layer that restricts water loss
Stoma
Epidermal pore regulating gas exchange and transpiration
Guard cells
Specialized cells controlling stomatal opening
Trichome
Epidermal outgrowth serving protective, secretory, or absorptive functions
Parenchyma
Living, relatively unspecialized cells involved in metabolism and storage
Collenchyma
Living support tissue with unevenly thickened primary walls
Sclerenchyma
Support tissue with thick, usually lignified secondary walls
Xylem
Vascular tissue transporting water and mineral nutrients
Tracheid
Elongated water conducting xylem cell with pits
Vessel element
Xylem cell joined end to end to form vessels
Phloem
Vascular tissue transporting sugars and other organic solutes
Sieve-tube element
Principal conducting cell of angiosperm phloem
Companion cell
Metabolically supports an associated sieve tube element
Plasmodesmata
Cytoplasmic channels connecting neighboring plant cells
Symplast
Interconnected cytoplasm linked by plasmodesmata
Apoplast
Cell walls and extracellular spaces outside the plasma membrane
Totipotency
Capacity of a cell to regenerate an entire plant
Protoplastid
Undifferentiated plastid that can develop into specialized plastid types
Chloroplast
Plastid responsible for photosynthesis
Amyloplast
Plastid specialized for starch synthesis and storage
Vacuole
Compartment involved in storage, degradation, tugor, and ion balance
Tonoplast
Membrane surrounding the vacuole
Primary cell wall
Flexible wall depositied while a cell is growing
Secondary cell wall
Additional wall deposited after cell expansion largely ends
Middle lamella
Pectin rich layer that binds neighboring cells
Cellulose
Beta 1,4 linked glucose polymer forming load-bearing microfibrils
Cellulose microfibril
Crystalline bundle of cellulose chains
Celluolse synthase complex
Plasma membrane complex that synthesizes cellulose
Cellulose synthase (CESA)
Catalytic protein responsible for cellulose chain formation
Hemicellulose
Wall polysaccharide that interacts with cellulose microfibrils
Xyloglucan
Major hemicellulose in many primary cell walls
Pectin
Hydrated wall polysaccharide important in adhesion and wall porosity
Homogalacturonan
Major pectic polymer composed principally of galacturonic acid
Lignin
Phenolic polymer that strengthens and waterproofs secondary walls
Suberin
Hydrophobic polymer deposited in cork and root barrier tissues
Callose
Beta 1,3 glucan deposited at plasmodesmata, sieve plates, and wound sites
Cell-wall matrix
Hemicelluloses, pectins, and proteins surrounding microfibrils
Wall loosening
Modification that permits irreversible cell wall extension
Expansin
Proein that promotes wall extension by weakening noncovalent wall interactions
Acid-growth hypothesis
Model in which proton extrusion activates wall loosening and expansion
Turgor pressure
Hydrostatic pressure that supplies the force for cell expansion
Cell-wall stress
Force per unit wall area produced by internal pressure
Cell-wall strain
Relative deformation of the wall under stress
Wall extensibility
Capacity of the wall to undergo irreversible expansion
Cortical microtubules
Cytoskeletal elements guiding cellulose deposition and growth direction
Anisotropic growth
Unequal expansion along different cellular axes
Oligosaccharins
Wall derived oligosaccharides that can act as signaling molecules
Water potential (Psi w)
Chemical potential expression predicting the direction of water movement
Solute potential (Psi s)
Reduction in water potential caused by dissolved solutes; also called osmotic potential
Pressure potential (Psi p)
Hydrostatic pressure contribution to water potential
Matric potential (Psi m)
Water potential effect of water binding to surfaces and matricies
Gravitational potential (Psi g)
Water potential component resulting from elevation
Chemical potential
Free energy per mole of a substance relative to a reference state
Osmosis
Movement of water accross a selectively permeable membrane
Osmotic pressure
Pressure required to prevent osmotic water movement
Van’t Hoff equation
Relationship used to estimate the osmotic potential of dilute solutions
Turgor presure
Positive pressure exerted by cell contents against the wall
Plasmolysis
Separation of the protoplast from the wall after water loss
Incipient plasmolysis
Point at which turgor pressure approaches zero
Water-potential gradient
Difference in water potential that drives water movement
Bulk flow
Coordinated movement of water and solutes caused by a pressure gradient
Diffusion
Net molecular movement for higher to lower chemical potential
Hydraulic conductivity
Ease with which water moves through a membrane or tissue
Aquaporin
Membrane channel protein facilitating water movement
Reflection coefficient
Measure of a membrane’s selectivity for a particular solute
Osmotic adjustment
Accumulation of solutes that helps maintain turgor under water stress
Pressure probe
Instrument used to measure cell turgor direclty
Psychrometer
Instrument that estimates water potential from vapor pressure measurements
Pressure chamber
Instrument commonly used to estimate leaf or xylem water potential
Relative water content
Tissue water content relative to its fully hydrated condition
Soil-plant-atmosphere continuum
Transpiration
Transpiration stream
Cohesion-tension theory
Cohesion
Adhesion
Surface tension
Capillarity
Negative pressure
Cavitation
Embolism
Root pressure
Guttation
Hydathode
Casparian strip
Endodermis
Exodermis
Apoplastic pathway
Symplastic pathway
Transmembrane pathway