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Seeds
Reproductive structures produced by angiosperms and other seed plants
Usually as the result of sexual reproduction
Contain embryos that develop into seedlings upon germination
Provide: Food
and Protective coating
Alternation of Generations
there is an alternation between a dipole (2N) plant form (Sporophyte) and haploid (1N) plant form (Gametophyte)
Gametophyte
Haploid, or gamete-producing plant form
Grow and develop within flowers of angiosperm
Produces gametes by mitosis
Sporophyte
diploid or spore-producing plant form
Large "plant" in flowering plants
Produces haploid spores by meiosis
Reduction division
2N to 1N
Syngamy
Fertilization
This process results in the formation of a diploid which undergoes mitosis to form an embryo (multicellular)
What is the embryo considered?
a sporophyte
What are the characteristics of the plant embryo?
a sporophyte that lies dormant in the seed with a supply of stored food and seed coats
What three parts is the plant body composed of?
stems, leaves and roots
What makes up the Shoot system?
stem and leaves
What makes up the Root system?
roots
Stem
produce leaves and branches and bear the reproductive structures
Leaves
flattened structures specialized for photosynthesis
Roots
provide anchorage in the soil and faster efficient uptake of water and minerals (storage)
Indeterminate Plant Growth
Increase in size as long as the plant is alive
What are the three means by which plants grow?
Increase number of cells -> cellular reproduction (mitosis and cytokinesis)
Increase in cell size
Increase in weight
What reproductive structures do mature plants produce?
flowers, seeds and fruits
Flowers are produced by what type of growth?
determinate growth
Flowers and floral buds are
reproductive shoots
Flower tissues enclose and protect
tiny male and female gametophytes
Fruits enclose seeds and function in
seed dispersal
meristem
region of undifferentiated cells that produce new tissues by cell division
Where are dormant meristems in seed embryos?
at root and shoot tips
Mature plants have
shoot apical meristems and root apical meristems
What powers the transformation of seedlings to mature plants?
Photosynthesis
What is the hierarchy of structure in a mature plant?
Specialized cells
Tissues
Organs
Organ systems (branches, buds, flowers, seeds, fruits)
Root and shoot systems
Plant (organism)
Primary Growth
- Elongation of plant organ(s)
- Roots, stems and leaves
- ALL plants
- Produces 6 Primary tissues
derived from apical meristems (RAM, SAM)
Primary xylem
vascular/conducting tissues ; water and minerals
Primary Phloem
vascular/conducting tissues ; food and solutes
Epidermis
dermal (outermost) tissue ; protection (1 cell thick)
Parenchyma
most abundant type ; storage ; part of cortex and pith
Collenchyma
protection/support of growing plant organs ; cortex
Sclerenchyma
protection/support of non-elongating organs ; cortex
Secondary Growth
- Expansion of plant organ(s) [lateral meristems]
- Roots and stems (NOT leaves)
- NOT all plants
- Produces secondary tissues (woody tissues)
What are the two major groups of angiosperms?
eudicots and monocots
Eudicots
- 240,000 species
- All with primary growth ; most with secondary growth
- Ex. Oaks, trees, roses, pansies
Monocots
- 60,000 species
- All with primary growth ; very few with secondary (we will consider it absent)
- Ex. Grasses, corn, tulips, lilies
What are the major functions of the root system?
- Absorbing water and minerals
- Anchoring the plant in soil
- Storing nutrients and water
Eudicots have what kind of roots
taproots
Monocots have what kind of roots
fibrous roots
What are the three regions of root growth?
- Region of Cell division
- Region of Elongation
- Region of Maturation
Region of Cell Division
(root apical meristem and root cap)
- RAM contains cells (divide and keep dividing)
- root tips embedded in lubricating mucigel
Region of Elongation
cells extend by water uptake
Region of Maturation
- root cell differentiation and tissue specialization
- Identified by presence of root hairs (water and mineral uptake) absent from older regions
Epidermis of mature roots
- encloses a cylinder of parenchyma called the root cortex
- Often rich in starch (function as a food storage site)
- May contain intercellular air spaces
Endodermis
selective absorption of minerals / (vascular cylinder)
Meristematic pericycle
- encloses root vascular tissue
- produces lateral (branch) roots
Woody roots produce
primary vascular tissues followed by secondary vascular tissues
What are the 4 parts of shoots?
Stem node, Internode, Leaf, Axillary meristems
Stem node
leaves emerge (or branch)
Internode
stem between adjacent nodes
Axillary meristems
- generate axillary buds
- Can produce flowers or branches (lateral shoot)
- New branches bear a SAM at their tips
- SAM (Shoot Apical Meristem)
Terminal bud
at end of each shoot; includes the SAM and other parts
Eudicots have what venation?
pinnate or palmate (NETTED)
- provide more support to the leaf
Monocots have what venation?
parallel
Stems
mostly above ground organs but some modified stems are below ground
Eudicot stems
- Primary (elongation) and secondary growth (expansion) in most
- Vascular bundles (xylem and phloem) form a 'ringed' pattern exhibit both a cortex and pith
Monocot stems
- Primary growth (elongation)
- Vascular bundles (xylem and phloem) are 'scattered' ; LACK both a pith and a cortex
Lateral Meristems
expansion
Lateral meristems produce
secondary tissues: = 2* Growth
What are the two lateral meristems?
vascular cambium and cork cambium
2 lateral meristems
both are 'rings' that retain cell division properties and produce 'rings' of secondary tissues to the inside/outside of cambium ring
Vascular Cambium
produces ring of secondary xylem (wood) to the inside and ring of secondary phloem (inner bark) to outside
Cork Cambium
produces rings of periderm (outer bark) that replaces the epidermis and cortex for external protection
Cork (outer bark)
periderm
Woody plants
- begin life with only primary vascular systems
- produce secondary vascular tissues and bark as they mature
Secondary xylem
wood
Secondary phloem
inner bark
Bark has both
outer bark (periderm) (mostly dead cork cells) and inner bark (secondary phloem)
Secondary growth in Eudicot stems
- Begins late in the 1st year of growth
- Eudicot stem after 3 years of secondary growth - 3 rings of 2* xylem (=wood)
Eudicot Roots
primary & secondary growth (most), cortex, no pith; core of xylem in the core of root
Monocot Roots
primary growth; cortex and pith
Hormones
- chemical messengers that regulate plant growth
- Most transported in phloem tissues
- ALL require an expenditure of energy on part of plant (ATP) for transport
- Interact with external environment factors to determine growth
Hormones control
Growth
Seed germination
Flowering
Fruiting
Shedding of leaves
Color change
Senescence
degradation of chlorophyll ; color change
What are the two broad categories of hormones?
growth inhibiting and growth promoting
Auxins
- First group of plant hormones described
- Growth promoting hormone
- Produced in:
- Shoot tips, seeds, fruits, leaves, stems (NOT roots)
Auxin effects
Promote cell elongation
Promote shoot elongation
Promote production of wood (2* xylem)
Promote fruit development
Inhibits lateral bud development (axillary buds)
Inhibits abscission (dropping of) of leaves, flowers, fruits
Cytokinins
- Originally detected in coconut 'milk'
- Growth promoting hormone
- Produced in:
- Seeds, fruits, roots
Cytokinin Effects
Promotes cell division (name derived from cytokinesis)
Promotes lateral bud development
Inhibits leaf senescence (change of color due to breakdown of pigments)
Gibberellins (gibberellic acid)
- Many types (more than any other group)
- Growth promoting hormone
- Found throughout plant but concentrated in seeds
Inside embryonic plant in seed
Gibberellins Effects
Promotes stem elongation by stimulating cell division and elongation
Promotes breakdown of food (starch) reserves in germinating seeds
Steps with gibberellin release
- Intake of water causes swelling and embryo hydration
- Embryo secretes gibberellins
- Gibberellins transported to cells of aleurone layer to secrete enzyme
(alpha-amylose) for breakdown of endosperm to glucose
- Embryo will respire glucose to produce ATP
- Embryo is directing the timing of germination
- ADVANTAGE of SEED PLANTS
Brassinosteriods
Growth promoting hormone
Brassinosteriod Effects
Promotes cell expansion
Promotes shoot elongation
Promotes shoot elongation
Promotes xylem tissue development
Promotes stress response
Inhibits lead abscission (dropping)
Abscisic Acid (ABA)
Growth inhibiting hormone
Found in large quantities in seeds moisture leaves, and dormant buds
Abscisic Acid Effects
Inhibits cell elongation
Inhibits alpha-amylase production
Promotes leaf senescence
Promotes production of storage proteins in seeds
Ethylene
Growth inhibiting hormone
Actually a gas produced by incomplete metabolism
Ethylene Effects
Promote fruit ripening
Promote abscission of leaves, fruits, flowers
Interacts with 4 growth promoting hormones to determine cell size/shape
Seed germination
Requires breaking of dormancy- combination of internal and external factors (environmental conditions)
Internal development of seedlings
Cells -> tissues -> organs -> organism
Nutrient
substance metabolized by or incorporated into an organism
What does photosynthesis require?
CO2, water, and elements such as potassium, nitrogen, and calcium
All but what is primarily taken up from the soil?
CO2
Deficiency symptoms
develop in plants that receive too little of a nutrient
Essential elements
play many roles in plant metabolism, often functioning as enzyme cofactors
Macronutrients
required in amounts of at least 1 g/kg of plant dry mass (>)
Micronutrients or trace elements
required in amounts at or less than 0.1 g/kg of plant dry mass (<)
Limiting factors
resources that can limit growth
Light
Carbon dioxide
Water
other mineral nutrients
13 soil nutrients
absorbed dissolved in H2O through roots and all follow the same pathway through plant as H20 (XYLEM)
Nitrogen (N)
macronutrient
component of proteins, nucleic acids, chlorophyll