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Evolution
Descent with modification, or change in the genetic composition, form and/or behavior of organisms over generations.
Two major contributors to evolution
Charles Darwin and Alfred Russel Wallace
2 major themes of biological evolution
Process (Natural selection, genetic drift, etc.) and history (what has evolved)
Age of Earth (Creationist perspective)
6000 years
Creationist view of species
do not change, created independently
Plato view of species
Species are unchanging types, variations unimportant, even misleading
Essentialism
Essential qualities or characteristics of things
Typological thinking
Putting things into categories based on characteristics
Great Chain of Being
Aristole, Scala Naturae, Fixed sequence based on increasing size and complexity, humans at top
1700s popular thinking
1. Species are fixed types
2. Some are higher- more complex or "better'
What changed the Pre-Evolutionary world view?
1. Challenges to the account of human history
2. Challenges to origin of the earth (geology, fossils, etc.)
Ussher
Calculated age of the earth based on Hebrew Patriarchs
Charles Lyell
Popularized James Hutton's concept of uniformitarianism
Uniformitarianism
that the Earth was shaped by gradual processes that are still active today
Charles Lyell's view of Species
Extinction yes, evolution no
Lamarck's Theory of Evolution
1. Organisms originated by spontaneous generation, evolve by moving up
2. Inheritance of acquired of characters. Change in response to environment passed on to offspring.
Assumptions of Lamarck's Theory
-Lower organisms show up
-To have evolution, things evolve from lower organisms to higher organisms
-Changing yourself can change offspring (sort of epigenetics)
Lamarck's view of Species
Species change through time
-Multiple origins of life
-No extinction (1809 before extinction became popular)
Buffon's Law
despite similar environments, different areas of the globe have different animals
Four Postulates of Natural Selection
1. Variation
2. Inheritance
3. Tendency towards population growth
4. Differential survival and reproductive success
Adaptation
heritable trait that increases fitness of an individual with that trait, compared to individuals without that trait, in a particular environment
Unit of natural selection
phenotype of individuals
Unit of evolution
population
Important points of natural selection
1. Mutation is random
2. Variability exists but heritable variability is only acted on by natural selection
3. New heritable traits results from mutations
Normal Curve
Bell-shaped distribution pattern
Narrow normal Curve
low variation
Wide normal Curve
high variation
Directional Selection
Change in average in a certain direction
Mechanisms of Evolution
1. Natural Selection
2. Genetic Drift
3. Sexual Selection
Stabilize Selection
Maintains intermediates phenotypes, reduces variation
Disruptive Selection
Intermediates selected against, maintains or increases variation, does not change mean and may lead to speciation
Hardy-Weinberg Principle holds true under...
1. Random mating
2. No natural selection
3. No genetic drift
4. No gene flow
5. No mutations
Genetic Drift
Allele frequencies drift randomly over time not adaptively
-Eventually leads to random loss or fixation of alleles
-especially pronounced in small populations
Founder Event
Group starts new population in new area
Founder effect
Allele frequencies differ from the original population
Genetic Bottleneck
Population becomes very small, and sudden reduction in alleles
Gene Flow
Movement of alleles between populations; individuals leave a population; join another and breed.
Sexual Selection
Animals evolved extravagant characteristics, colors and displays to better obtain mates and increase reproductive success (traits aren't always adaptive in the context of natural selection)
Parental Investment
Whatever parent does to increase survival of existing offspring at the cost of the parent's ability to generate more offspring.
Limiting Factors for sexual selection for females
-Her ability to produce eggs
-Ability to provide parental investment
Limiting factors for sexual selection for males
-Ability to secure mates
-Competition with other males
-Females by choice
Species
An evolutionary independent population or group of populations
3 Approaches to identify species
1. Biological Species Concept
2. Morphological Species Concept
3. Phylogenetic Species Concept
Creator of Biological Species Concept
Ernst Mayr
Biological Species Concept
Species are defined based on reproductive isolation (cannot interbreed or fail to produce viable offspring)
Disadvantages of Biological species concept
-Cannot be evaluated in fossils it in species that reproduce asexually
-Can only be applied to populations that overlap geographically
Prezygotic Isolation
Occurs when there is a barrier to stop mating
Types of Prezygotic Isolation
1. Temporal- breed at different times of a year
2. Habitat (physical isolation)
3. Behavioral
4. Gametic (sperm and egg are not compatible)
5. Mechanical (parts don't fit)
Postzygotic Isolation
Produce hybrid with low fitness, hybrid in-viability, or hybrid is sterile
Morphospecies Concept
Species are identified be difference in morphological features (traits arise from isolation)
Disadvantages of morphospecies concept
-cannot identify a cryptic species
-features used to distinguish are subjective
Phylogenetic Species Concept
Defined by cladistic monophyly
Speciation
Splitting of populations of a single species due to genetic isolation
Allopatric Speciation
Speciation caused by physical isolation and genetic isolation
Dispersal
Mode of allopatric speciation- colonizes new habitat forms new population
Vicariance
mode of allopatric speciation- physical barrier, isolated subgroups
Sympatric Speciation
Speciation in same geographic area
Polypoidy speciation
Type of sympatric speciation- where gametes don't split chromosomes correctly, common in plants
Polytomy
More than one branch from one mode
Homology
Similarity due to a shared ancestor
Homoplasy
similarity be convergent evolution
Synapomorphy
Shared derived character state
Symplesiomorphy
shared primitive character state
Apomorphy
uniquely derived characters that are specific to just one taxon
Monphyletic group
All organisms on a branch of phylogenetic tree descended from a single common ancestor