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How do bird wings produce lift?
A bird’s wing is shaped to be bent
Cambered wing
More lift is generated with higher air speed
Cambered wing
Causes air to move faster on the top of the wing than the bottom of the wing
Induced drag
A cost of lift. Happenes when there are pressure differences in the upper and lower surfaces of the wing.
Can be reduced with wide spaces in outer primaries
Reduced with pointier wings
Strongest at slow speed
Alula feathers
Reduce drag. Direct the air to follow the wing more closely, leading to less turbulence.
Profile drag
The drag produced by the profile of the bird as it moves through the air (the bird “displacing” the air)
Increases with air speed
How ro reduce profile drag?
Bird’s aerodynamic shape and tail
Long and narrow wings
Minimum power speed
The bird is able to stay in the air long, but not maintain a high speed
Maximum range speed
Gives the fastest range with the least energetic cost
What forces affect a bird in flight?
Upward flight: Lift, offset by weight of bird and gravity
Downwards flight: Drag, offset by wind+ updrafts, wing flapping
How can a bird offset the downward forces it experiences in flight?
Downwards flight: Drag, offset by wind+ updrafts, wing flapping
What external forces can birds use for flight?
Thermal soaring- rising in warm air
Slope soaring- rising in mountain updrafts
Dynamic soaring- Flying into a strengthening wind and with a weakening wind
Flapping flight
Requires enough power per body mass of a birds to maintain flight. Eventually birds that are too big do not have enough power to paintain flight (power decreases as body mass increases
Explain the Power/Velocity plot
Larger species have limited power, which changes the flight style of the bird
Hummingbirds have a small body size, and can produce lots of power (more than needed) for hovering
Rock pigeons are larger, and can hover briefly
Vultures need to run to get enough power needed to fly
The condor needs wind and thermal currents to be able to fly
Why do some birds fly in formations?
The bird flying in front provides upward turbulence to the birds behind
Intermittent flight
Flapping phases are interrupted by pauses between each
May include:
Bounding- wings folded next to body (small to medium birds)
Gliding- wings stretched out (medium to large birds)
Wing loading
The bird’s body mass per wing area (higher loads require fast speeds & manaeuverability)
Aspect ratio
The wingspan squared divided by the wing area (high aspect ratio (often) = pointy wings)
What different types and subtypes of movement do you know?
Local movements
Residence
Dispersal (natal dispersal + breeding dispersal)
Philopatry
Migration (facultative, obligate, partial)
Nomadism
Irruption
Local movements
Short trips (e.g. for foraging)
Residence
Same location all year
Dispersal
Movement between breeding sites
Natal dispersal
Moving from hatch site to first breeding site
Breeding dispersal
Moving to a new breeding site in the next breeding season
Philopatry
Returning to the same location for several years
Migration
Seasonal movements between breeding and wintering sites across latitudes or altitudes.
Facultative migration
migration pattern and timing changes according to the environment
Obligate migration
Migration that follows the same pattern and time every year
Partial migration
A species that includes individuals that migrate and those that don’t (residents)
Nomadism
Irregular movements to breed where resources are best
Irruption
Birds that occasionally move in large numbers beyond their migration range in response to poor food supplies (facultative and nomad)
Why do birds migrate?
To get seasonal resources elsewhere
To avoid harsh conditions (low food, light, cold)
Fewer parasites & diseases (in winter areas)
Benefits of resources outweigh csts of migration
Loop migration
A species that takes a different route north and south
Leapfrog migration
North species breed north, and south species breed south
What migration strategies do birds use?
Long-distance migrant: Magnolia warbler, American Redstart
Loop migrant: Blackpoll warbler
Leapfrog migrant: Fox sparrow
Broad front migration
A pattern in which the migrating individuals of a species are spread across a wide area rather than narrower flyways.
Soaring migration
Use thermals to soar during the day (avoid water)
Hop
Short bouts of migration and fueling stops (fly at night, rest during day)
Skip
Long flights with long refuling periods (large unsuitable habitats)
Jump
Very long non-stop flight (requires lots of feeding prior) (Bar-tailed godwit)
What makes shorebirds vulnerable during migration, and what are threats to them?
Shore birds rely on a few high-quality sites (it effects their survival, thus needs conservation)
Threats of shore birds:
Harbors & airports
Shrimp ponds and aquaculture
Oil spils
Climate change
Hunting disturbance
How can we study bird migration? What are some advantages and disadvantages of the different methods to study bird migration?
Banding
Visual counters
Geolocatiors: Log daylight & require recapture (covered nests may fool lights)
GPS loggers/ satellite trackers for large birds
Feather isotopes: Shows where a feather was grown
Genetic information: origins of migrants
Experiments: Displacement & manipulation
Radars
How do birds chose when to migrate and at what altitude?
Tradewinds
Tradewinds
Wind changes direction at cerain altitudes and at cerain times of the year
Weather radars
Can be calibrated with specialized bird radars & sophisticated computer software can be used to track bird migration at a continent-wide scale
Uses: Shows weather influences, monitored in real time, migration forecasts
What cues may birds use for orientation and how do they use them?
Annual cycles
Magnetic perception
Solar orientation
Celestial navigation
Odor
Infrasound
Landmarks
Learned & innate orientation
Magnetic preception
Birds have a pigment cryptochrome in their eye that may allow them to see magnetic inclination in presence of blue light
Shows north south poles, and east and west
What experiments help to study bird migration and the use of specific cues?
Adjusting perception of light
Changing stars visible (solarium)
Magnet on birds- to upset magnetic orientation
Displacement
Emlen funnel experiments
Migratory divide
Locations where two populations of the same species with different migratory orientations meet and potentially interbreed.
What information would you need to collect in a specific case to show that a migratory divide contributes to speciation?
The origin of the birds
Migratory routes
Hybrids with intermediate migratory behavior
Selection against hybrids
Ecology
The study of interactions of organisms with each other and the environment
Abiotic factors
Physical features of the environment (e.g. temp., topography, rainfall, altitude)
Biotic factors
Features of the environment that result from other living organisms (cometition, predation, food)
Population
Individuals of the same species living in the same area at the same time
Density-independent factors
Stuff outside that may affect individuals despite their density (no benefit to certain densities e.g. natural disasters)
Density-dependent factors
There are costs/ benefits to having certain densities (e.g. parasitism/food availability)
Calculating pop. size changes
Birth - Death + Immigration - Immigration
Influences on pop. Size
Mortality
Sex ratio
Competition
Predation
Food
Weather
Brood parasites
Pathogens
Nest site availability
Trade-offs
Two directions: one in which they produce many offspring and have low survivorship, and the other in which they produce few offspring and have high survivorship
R selection
many offspring and low survivorship (small birds)
K selection
few offspring and high survivorship (large birds)
More susceptible to mort
Density dependent regulation
As the density in a population increases, the amount of offspring they produce (fecundity) decreases. If there is a very low density, offspring production also decreases.
Allee effect
- a tendency for populations at low density to have poorer, rather than better, reproductive success (falls below a threshold level)
Causes of allee effect
Difficulty finding mates
Lack of group protection from predators
Lack of group cooperation
Inbreeding
Uneven sex ratios
What are your take-aways from the discussion of the journal article by Rodenhouse et al.?
Ways to regulate population size:
Interactions among individuals (crowding, territoriality)
Environmental heterogeneity (variation in environment)
Intrinsic differences among individuals
Methods
Warbler density was manipulated- One group of territories were left alone, and in a few random territories all conspecifics were removed
As population size grows, most birds choose to go to the best sites. Once these sites are filles, warblers will have to settle for lesser quality sites (could be lesser quality due to crowding)
Community
Interacting organisms of different species
Assemblage
A particular subset of a community
Ecosystem
Biological community plus physical environment
Niche
Biotic & abiotic factors that allow a species to occuR
Fundamental niche
Possible niche based on environmental factors
Generalists
Species with broad niche breadth
Realized niche
Niche that is actually occupied by a specieS
Specialists
Species with narrow niche breadth
Density inflation
an increase in population size when limiting factors (such as competitors or predators) are removed
Character displacement
when species evolve differences to reduce competition when they coexist, leading to greater specialization.
Density compensation
when some species become more abundant to compensate for the loss of others, maintaining overall population density.
Ecological release
when a species expands its niche due to reduced competition, often using a wider range of resources or habitats.
Species diversity
Number of a species
Species richness
includes both species richness and species evenness (the relative abundance of each species)
What hypotheses explain the high species richness in the tropics?
Higher speciation
Lower extinction rates
Complex habitat structure & less seasonal
What are problematic causes of mortality for birds of prey?
Eating poisoned rodents → Don’t use rodenticides
Lead poisoning from eating lead bullets → Don’t use lead bullets
Mercury poisoning from eating fish → Don’t use/release mercury from factories
Electrocution from power lines → Good construction
Death from wind farms → Better placement of turbines
Car collisions → no food scraps on roadsides
What are major causes of the Anthropocene mass extinction?
Habitat modification, fragmentation, destruction
Overexploitation
Invasive species
Pollution
Climate change
What are examples of human caused extinctions of birds?
Dodo- Over hunting
Great Auk- Over hunting
Guam rail and Guam kingfisher- Invasive species introduction
Why are island species particularly vulnerable, and what are some of their main threats?
Sometimes not able to fly away from predators
Habitat loss
Inbreeding → decreased genetic variation
Not as disease resistant
What was the likely most abundant bird species to have ever lived, and why did it go extinct?
Passenger pigeon- Overhunting/ destroyed habitat