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a species
is a group of similar organisms that can interbreed in nature and produce vigorous, fertile offspring
subspecies are
different races of the same species, who do not interbreed directly (gene flow can still exist)
extinction
the permanent loss of a species (usually hasn’t been seen for 50 years or more)
a population is a
group of organisms of the same species living in a defined geographic region at the same time.
gene pool:
the genetic makeup of a population; includes the sum of all the alleles for different genes present in a population
gene flow is the
movement of genes and alleles between populations
gene flow can occur through
immigration or emigration or random mating between individuals
Immigration
= individuals move into a population → bring in new alleles
Emigration
= individuals move out of a population → remove alleles.)
gene flow increases the
genetic diversity within a population
when gene flow stops
speciation can occur
variation is important for a
population to survive any sudden change in their environment
variation occurs when
members of a population display different phenotypes (traits)
variation traits can include
physiological, structural, or behavioural
sources of variation are (reg)
reproduction
environment
gene mutation
reproduction source of variation
sexual reproduction (lots)
asexual reproduction (little)
gene mutation variation
change in DNA base sequence
introduces new alleles, leading to new genetic variation
only passed on if mutation occurs in germline cell
environment variation
acts as a selection pressure on phenotypes
high genetic diversity in a population
large population size
random mating by sexual reproduction
immigration
mutations resulting in different phenotypes
allele frequencies can measure
genetic diversity by examining the gene pool of a population and calculating the allele frequency
allele frequency differences may be due to chance or due to selection by selection pressures
if a population has a wide range of alleles in their population it means they are genetically diverse
allele frequencies
how common an allele is in a population is referred to as the allele frequency
does the allele frequency in each population stay the same?
no the allele frequency in each population will differ
allele frequency differences may be due to
chance or due to selection by selection pressures
if a population has a wide range of alleles in their population it means
they are genetically diverse
hardy weinberg equilibrium allele frequency formula
p+q=1
p: represents the frequency of the dominant allele.
q: represents the frequency of the recessive allele.
hardy weinberg equilibrium genotype frequencies formula
p² + 2pq + q² = 1
p: represents the frequency of the dominant allele.G
q: represents the frequency of the recessive allele.g
p²: represents the frequency of the homozygous dominant genotype. GG
2pq: represents the frequency of the heterozygous genotype.Gg
q²: represents the frequency of the homozygous recessive genotype.gg
q =
total recessive alleles/total alleles in population
p =
1 - q
genetic drift refers to
the random changes in allele frequency due to chance events
what populations does genetic drift occur in
occurs in all populations
what populations does genetic drift have a more pronounced effect on and what can it result in
it has more of a pronounced effect in small populations and can result in the loss or fixation of an allele
can result in reduced genetic diversity in small populations
types of genetic drift
bottleneck effect, founder effect
the founder effect occurs when .. and results in a
a few individuals migrate to create a new population in a different location that is not representative of the original population
the individuals are genetically different to the original population resulting in a different gene pool
could occur due to habitat fragmentation (where a population becomes separated)
the founder effect often results in … due to ..
often low levels of genetic variation due to only a few founding individuals
the bottleneck effect
the severe reduction of individuals in a population generally due to a catastrophic event (e.g. disease, bushfire, human impact)
the bottleneck effect over time population will increase in numbers but
the genetic variation stays low as only have the gene pool from surviving individuals
geological timescale
the oldest fossils are between 3 billion and 3.5 billion years old. these are fossil bacteria, and for most of earth hisstory, life was simple.
more complex animals appeared in the oceans about 565 million years ago and became much more common about 542 mullion years ago
this last section in time is the start of a division of geological time called the phanerozoic eon. phanerozoic means “visible life” and is the time in which fossils are abundant
fossil succession
fossils appear in an order of “fossil succession” from single cellular organisms to structurally complex multicellular organisms.
significance of mass extinctions
several mass extinctions have occurred throughout earths history
mass extinctions provide opportunities for other species to thrive and diversify
fossil record
the information derived from fossils. The fossil record is arranged in chronological order and helps us map the history of life on Earth, placing species in the appropriate geologic time frame
fossils
the remains of an organism or direct evidence of its presence once on the earth
fossil record is incomplete (there are missing species in the evolutionary chain) as
most organisms do not get fossilized (e.g. organisms with hard body parts are more likely to fossilize than those with soft parts)
or the fossils are too old and there is no DNA (which means we cant determine how the fossil is related to other species)
most fossils are found in
sedimentary rock
how are fossils formed in sedimentary rocks
formed by mineralization
body parts are replaced by hard minerals as the surrounding sediments turn into rock
conditions for fossilisation (RUD)
-very rare
rapid burial (R)
covered in sediment (usually under water)
organism lies undisturbed (U)
no predators
no earthquakes, etc.
decomposition is prevented (D)
low o2, low temp, etc, to prevent bacterial decomposition
types of fossils
body fossil
trace fossil
index fossil
transitional fossils
body fossil
fossils of the actual organism
e.g. bones, teeth, and claws
soft tissues (e.g. tissue, muscle, organs etc) are very rarely fossilized
trace fossil
not the actual organism
e.g. burrow, footprints, faeces
index fossil
a fossil that:
had a wide geographic distribution
existed for a relatively brief period of time
fossils that are found in the same rock layer can be estimated to be the same age
a form of relative dating
transitional fossils
intermediate forms of organisms that provide an evolutionary link between species
e.g. archaeopteryx shows features of reptiles and birds
dating techniques
two methods are used to provide an age (date) for a fossil
relative dating
absolute (radiometric) dating
relative dating
the age of one rock is determined by comparing it to another rock
a fossils relative age is provided
uses rock layers (formed by stratification) to assign comparative ages to fossils
relative dating is based on. and uses what
the law of fossil succession and uses index fossils
stratigraphy/stratification
the study of the sequence of rock layers (strata) in any one area
layers deposit over time
older layers are lower
index fossils relative dating
fossils that can be found in the same rock layer as an index fossil can be estimated to be the same age
the principles of correlations allow the
relative age of rocks in different parts of the world to be inferred, based on distinctive index fossils
Occasionally, a strata representing a particular time period may be missing due to
environmental conditions (e.g. flooding, erosion, etc.)
relative dating definition
a dating technique used to determine the relative age of a fossil by comparing its position to other fossils or rock in surrounding rock strata (layers)
absolute (radiometric) dating
provides a numerical (absolute) age for a fossil
the relative amount of parent/daughter is compared
half life -
the time taken for half of the parent isotope to decay into its daughter isotope
absolute (radiometric) dating based on
radioactive isotopes and their half lives
isotope carbon-14 decays to .. half life ~.. used to determine..
nitrogen-14,
has a half-life of approximately 5,700 years,
the age of deposits less than 50,000 years old.
isotope uranium-235 decays to .. half life ~.. used to determine..
lead-207,
has a half-life of about 704 million years,
the age of deposits older than 500,000 years.
isotope potassium-40 decays to .. half life ~.. used to determine..
argon-40,
half-life of around 1.3 billion years,
the age of deposits older than 500,000 years.
carbon dating
date organic (contain C) fossils, less than 50,000 years
half life is approx. 5,700 years
fossilisation
: Organism is buried in sediment or water, gently covering the body.
Preservation conditions:
Minimal disturbances (scavengers, fungi, bacteria, earthquakes, uplift) to keep the fossil complete or nearly complete.
Low oxygen (anaerobic) environment slows decomposition, preserving bones (soft tissues may still decompose).
Discovery: Fossils are often uncovered later due to earth movements.
selection of phenotypes
some individuals are more likely to survive change in their environment due to a particular phenotype
at a selective advantage when the selection pressure is applied
genetically fit
Organisms with a more favourable phenotype are at
an adv as more likely to survive, reproduce and pass on alleles to their offspring
e.g. antibiotic resistance in bacteria
selection pressures
predators
availability of resources (e.g. water, shelter, mating habitat. etc) → due to competition between individuals within a population
food sources → due to competition
disease/pathogenic spread
environmental phenomena (e.g. natural disasters)
weather conditions/changes (e.g. floods, storms etc)
abiotic conditions (e.g. temperature, co2 concentration)
natural selection
proposed by darwin (and wallace) in 1860s
VIS: Variation in population Selective pressures Inheritance
variation exists in a population due to mutation
individuals who have a favourable adaptation/trait will be better suited to the environmental selection pressures
these individuals will survive and reproduce to pass on their favourable characteristics to their offspring resulting in changes to the allele frequencies
natural selection vs lamarckism
lamarckism
proposed that trait develops in response to the environment
use and disuse occurs (if u want it it will happen)
the process of natural selection
variation exists in individuals within a species that may lead to an advantage if a change in the environment occurs
variation exists in the population due to mutations
a selection pressure is present in the environment
this means some individuals in the population have a survival and reproduction advantage
individuals that survive pass on the allele to the next generation (through reproduction), resulting in more individuals with the trait
this results in increased allele frequencies for the trait in the population
selective breeding
uses the principle of natural selection
a human (usually a farmer) selects the trait that is desired based on phenotype
selects two organisms that are showing the trait and breeds them (to increase the chance of offspring having these traits)
observes the offspring and then selects the offspring with the best version of the desired trait. breeds the offspring
completes the process many times until the trait is observed in all offspring
note that the trait might be favourable to humans but not necessarily to the organism as it may reduce genetic diversity in the population
speciation process
there is variation in the population. there is a lack of reproduction between individuals in the population preventing gene flow (may be due geographic barrier or prezygotic barrier resulting in reproductive isolation)
different selection pressure act upon each population due to different niches
over time many different mutations occur in the different populations and they accumulate resulting in phenotypic differences
once significant differences have accumulated (and the species can no longer interbreed) they are considered to be different species
speciation def
process by which new species are formed
a species is a group of very similar organisms that can interbreed in nature and produce vigorous, fertile, offspring.
Pre-zygotic Isolating Mechanisms
Temporal
Ecological
Behavioural
Mechanical
Occurs when physical differences prevent copulation / pollination
Example: Certain breeds of dog are morphologically incapable of mating due to size
Pre-zygotic Isolating Mechanisms Temporal
Occurs when two species mate at different times of year
Example: Frogs live in same pond but breed during different seasons (summer vs spring)
Pre-zygotic Isolating Mechanisms Ecological
Occurs when two species occupy different habitats
Example: Lions and tigers can potentially interbreed, but usually occupy different habitats
Pre-zygotic Isolating Mechanisms Behavioural
Occurs when two species have different courtship behaviours
Example: Certain groups of birds will only respond to species-specific mating calls
Pre-zygotic Isolating Mechanisms Mechanical
Occurs when physical differences prevent copulation / pollination
Example: Certain breeds of dog are morphologically incapable of mating due to size
Post-zygotic Isolating Mechanisms
Hybrid Inviability
Hybrid Infertility
Hybrid Breakdown
Post-zygotic Isolating Mechanisms Hybrid Inviability
Hybrids are produced but fail to develop to reproductive maturity
Example: Certain types of frogs form hybrid tadpoles that die before they can become a frog
Post-zygotic Isolating Mechanisms Hybrid Infertility
Hybrids fail to produce functional gametes (sterility)
Example: Mules are sterile hybrids resulting from mating between a horse and a donkey
Post-zygotic Isolating Mechanisms Hybrid Breakdown
F₁ hybrids are fertile, but F₂ generation fails to develop properly
Example: The offspring of hybrid copepods have less potential for survival or reproduction
Allopatric speciation relies on a
geographical barrier to reproductively isolate groups of organisms in a species, leading to the formation of a new species.
allopatric speciation and steps
speciation due to geographic separation
there is variation and gene flow in a population living in a specific geographic area
a geographic barrier separates the population (e.g. a river forms or a mountain range forms) and gene flow stops
different selection pressures on either side of barrier. different phenotypes are selected for on either side
over time many different mutations occur in the different populations, and they accumulate resulting in phenotypic differences
the two populations become so different that they become reproductively isolated. if the two populations were brought back together, they could not interbreed. a new species have formed
allopatric speciation - galapagos finches
ancestral finch arrived from mainland south america
finches increased in number and the population many were adapted to the environment
some finches flew to other islands with different selection pressures (food source)
over time, the beak shapes changed and the finches became so different on each island that they became different species
sympatric speciation steps
there is variation and gene flow in a population living in a specific geographic area
a reproductive barrier prevents some members of the population from reproducing (e.g. mating at different times of the year)
over time many different mutations occur in the different populations, and they accumulate resulting in phenotypic differences
the two populations become so different that they become reproductively isolated. although the populations live in the same niche, they cannot interbreed. a new species has formed
sympatric speciation howea palms - lord howe island
one of the worlds best examples of sympatric speciation, when two species from a single ancestor form in the absence of a geographical barrier
the two species, the kentia or thatch palm (howea forsteriana) and curly palm (h belmoreana), appear to have diverged after they began flowering at different times of year, as a result of differing soil preferences
Sympatric speciation involves the
formation of a new species in populations located in the same geographical location
structural morphology
studies the sizes, shapes and structures of plants, animals and microbes, as well as the interactions between their parts
homologous structures:
have the same structure but may have a different function
evidence of different selection pressures
evidence of a common ancestor (divergent evolution)
analogous structures:
a different structure but the same function
due to similar selection pressures
no common ancestor (evidence of convergent evolution)
vestigial structures
a vestigial structure is a feature that a species inherited from an ancestor but that is now less elaborate and functional than in the common ancestor
due to a lack of selection pressure
vestigial structures example
e.g. whales still have a pelvic bone, however it is much smaller in size due to a lack of selection pressure (e.g. live in water vs live on land)
e.g. flightless birds like emus/ostriches have reduced wing sizes, due to lack of selection pressure (e.g. food sources on the ground)
molecular homology
study of the similarity or difference or molecular information (nucleic acid or amino acid sequence)
can determine evolutionary relationships based on how similar/different the sequences are
often used to construct phylogenetic trees
when talking about how related are species are acronym
TAN (time, ancestor mutations)
very closely related →
less mutations as shorter time of divergence
molecular homology
study of the similarity or difference of molecular information (nucleic acid or amino acid sequence)
nucleic acids can be analysed and compared for differences
includes nuclear DNA, mRNA, rRNA, mitochondrial DNA
chloroplast DNA