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Structures for hearing
Cochlea: snail-shaped chamber of inner ear
Modiolus: bony axis of the spiral
Cochlear duct: membranous labyrinth in cochlea. Roof formed by vestibular membrane. Floor formed by basilar membrane
Scala vestibuli: chamber of bony labyrinth adjacent to vestibular membrane
Scala tympani: chamber of bony labyrinth adjacent to basilar membrane
Helicotrema: small channel connecting scala vestibuli and scala tympani
Spiral organ: sensory structure for hearing. Within cochlear duct. Thick sensory epithelium
consisting of hair cells and supporting cells on basilar membrane
Hair cells: receptors that release neurotransmitter to sensory neurons.
Single row of inner hair cells
three rows of outer hair cells
Hair cells have many stereocilia (long microvilli) and one kinocilium (long cilium) at their apex
Stereocilia and kinocilium embedded in tectorial membrane.
Base of hair cells synapse with sensory neurons.
Cell bodies of sensory neurons are located in spiral ganglia of
modiolus
Pathway from sound wave to nerve signal
waves vibrate tympanic membrane
ossicles vibrate and transmit waves to oval window
fluid pressure waves in scala vestibuli push vestibular membrane causing pressure waves in endolymph of cochlear duct
specific regions of basilar membrane move
hair cells distorted causing changes in NT release
sensory neurons in CN VIII stimulate to fire
pressure transmitted to scala tympani
absorbed in round window
Cochlear hair cell stimulation
basilar membrane moves up → hair cells pushed into tectorial membrane
hair cells release NT from base → excite sensory neuron → action potential
basilar membrane down → reverses process
Perception of sound
sound = pressure waves from vibration
pitch = frequency
frequency = rate of vibration (Hz)
humans hear 20-20,00 Hz, sesnstive to 1500 and 4000
loudness = wave amplitude
Deafness
conductive: inference of wave transmission in external/middle ear
sensorineural: malfunction inner ear/cochlear nerve