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Ruminant stomach (IN ORDER!!!!)
4 compartments: rumen, reticulum, omasum, abomasum
What is the rumen also known as
paunch
rumen (paunch)
largest compartment
40-50 gallons in cattle
2.5-8 quarts in sheep
what does the rumen include
fermentation vat
microbes digest cellulose and starch
anaerobic bacterial fermentation
what does anaerobic mean
no oxygen
what is the reticulum also known as
honeycomb
reticulum (honeycomb)
sorts particles by size
sends large particles back for rumination
chewing cud
catches foreign objects
hardware stomach
what is the omasum also known as
many plies
omasum (many plies)
multiple muscular folds
mechanical grinding of feed
absorbs water, VFAs, electrolytes
what is the abomasum also known as
true stomach
abomasum (true stomach)
acidic
functions like monogastric stomach
produces HCl and enzymes (pepsin, rennin)
where does protein digestion begin
abomasum
rumen microbes and symbiosis
bacteria, protozoa, fungi work in symbiosis
digest cellulose and starch
synthesize microbial protein and vitamins
produce VFAs as energy source
fermentation products
volatile fatty acids (VFAs) and microbial protein
what are volatile fatty acids
main energy source (50-70%)
acetate (VFA)
from fiber fermentation → used for fat synthesis
propionate (VFA)
from starch and sugars → covered to glucose via liver
butyrate (VFA)
from fiber and starch → energy by rumen wall, fat synthesis
what is microbial protein
amino acids for growth and tissue repair
byproducts of fermentation
gas: methane and CO2 → belching (eructation)
heat production → rumen produces heat
essentrial for rumen function
monogastric stomach functions
storage and mixing of feed
secretes acid (HCl) and enzymes (pepsin)
begins protein digestion
what are the four regions of the monogastric stomach
esophageal, cardiac, fundic, pyloric
esophageal (monogastric)
no secretions, prone to ulcers
cardiac (monogastric)
secretes mucus to protect stomach lining
fundic (monogastric)
secretes HCl and enzymes (main digestion region)
pyloric (monogastric)
secretes mucus, controls passage to small intestine
enzyme action in monogastric: amylase
starch → sugar
enzyme action in monogastric: pepsin
proteins → peptides
enzyme action in monogastric: lipase
fats → fatty acids and glycerol
what does enzyme action in monogastric depend on
stomach pH
limitations and diet needs of monogastric
limited fiber
require nutrient-dense feeds (grains, protein meals)
balanced amino acids critical (can’t synthesize all)
what does growth efficiency depend on in monogastrics
feed quality
hindgut fermenter
horses, rabbits, and guinea pigs
use roughages because of active cecal bacterial population that digests fiber
hindgut fermenter: small stomach
only 8% of capacity in entire tract (3-4 gallons)
less muscular
feed arranges in layers
hindgut fermenter: large intestine
60% of total gut capacity
cecum and large colon contain bacterial population similar to ruminants
beak
prehension
the crop
expansion of esophagus
temporary feed storage
softens food before digestion
some microbial activity
proventricules
secretes HCl and pepsin → chemical digestion
gizzard
muscular, grinds feed with grit
small intestine: poultry
main site for nutrient absorption
two ceca
some fiber fermentation; limited
large intestine
very short, limited function
cloaca
single exit for feces, urine, eggs
poultry digestive system
very fast (2-4 hours)
nutrient-dense food required
cannot digest fiber efficiently
balanced amino acids essentail
feeds
delivery system for nutrients
the right feed choices impact:
cost efficiency, animal health, sustainability
nutrient composition
amount of specific nutrients, expressed as % of the dry matter
dry matter
portion of feed left after water is removed, contains nutrients
crude protein (CP)
measures nitrogen content of a feedstuff
digestibility
amount of absorbable nutrient in a feed
crude fiber (CF)
fiber contents in feeds
total digestible nutrients (TDN)
sum of digestible fiber, protein, lipid, and carbs components of a feedstuff
what value is most commonly used in simple ration balancing
Total digestible nutrients (TDN)
dry matter
the universal basis for comparison
why dry matter
water dilutes nutrient concentration
always compare feeds on a DM basis
forage testing reports are DM standardized
NRC feed classifications
dry forages and roughages
pasture and green forages
silages
concentrates (energy foods) (carbs)
protein supplements
mineral supplements
vitamin supplements
nonnutritive additives
right feed for the need
start with animal’s requirement, not the feed
identify which nutrient is limiting production
choose the feed class that supplies it most efficiently
right feed: energy
concentrate
right feed: protein
protein supplement
right feed: bulk/fiber
forage or roughage
right feed: vitamin/mineral
supplement/additive
dry forages and roughages
higher in cellulose, hemicellulose, lignin
lower energy density than other feeds (high in fiber)
required for rumen health, stimulates chewing and saliva
maturity at harvest
more mature = high fiber, low protein
weather damage
leaches nutrients
soil fertility
impacts proteins and mineral levels
plant species
legumes > grasses for protein
harvest method
mold, spoilage
pasture and green forages (seasonal nutrition)
most common global feed source
young, fertilized pasture = high protein, highly digestible
mature, overgrazed pasture = low protein, high fiber
rational grazing
allows rest and regrowth, maintains plant health
fertilization and soil health
improves protein content and yield
weed control
reduces competition for nutrients
avoid overgrazing
prevents pasture degradation
silage
preserving nutrients through fermentation
what is silage
anaerobic fermentation: beneficial bacteria lactobacillus produce lactic acid
preserves forage by decreasing pH
does not increase nutrients- fermentation consumes some energy
risk of mycotoxins if storage is poor
corn silage
most common, increase energy from starch
haylage
fermentation grass or alfalfa, increase protein
by-product silages
cannery waste, root crops, beet pulp
concentrate
high energy, high risk, low fiber
how much concentrate in crude fiber
< 18%
how much concentrate in crude protein
< 20%
concentrates
increase in starch → rapid energy release
risk digestive upset if fed to quickly or in excess
rumen acidosis
where do most concentrates come from
corn, barely, molasses
protein supplements
building blocks for growth
protein supplement in plants
SBM, cottonseed meal, peanut meal
protein supplement in animal
fish meal, blood meal, feather meal (cannot be fed to ruminants)
protein supplement in nonprotein nitrogen (NPN)
urea - ruminants only!
nonnutritive additives - beyond nutrition
growth promo: ionophores, hormones
health: anti- and pro- biotics
palatability: flavors, coloring agents
controlled by FDA and USDA
Modern feeding challenges: sustainability focus
reduce methane emissions
decrease carbon footprint of feed production
Modern feeding challenges: precision feeding
AI-driven systems to tailor diets to individual animals
Modern feeding challenges: alternative proteins
insects, algae, fermentation-based feeds
why ration balancing matters
prevents deficiencies → poor health and reproduction
avoids excesses → waster and cost
meets animals performance goals efficiently
Pearson Square
developed in late 1800s
1st simple visual method for balancing livestock rations
Pearson Square- simple configurations
balances 2 feeds to reach a target nutrient level
works for protein, energy, or minerals
goal value must be between 2 feed values
impact of farm animal reproduction
essential for species survival and livestock production
primary driver for profitability in animal agriculture
efficiency gains in repro increase impact than most other measures
3% increase in birthrate yields
~ +1 million beef calves/year
~ +900K pigs/year
~ +6.8 billion gallons of milk/year
endocrinology
study of glands and hormones
main control center
reproduction controlled by Hypothalamic-Pituitary-Gonadal (HPG) Axis
central neuroendocrine system
brain and reproductive organs must coordinate
why neuroendocrine control matters
nutrition and body condition
stress and health
photoperiod (day length)
hypothalamus = headquarters
brain region controlling homeostasis
physiological and behavioral functions
appetite and energy balance, body temp, stress responses, repro
What does the hypothalamus release
GnRH (Gonadotrophin-Releasing Hormone → signal pituitary