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evolution definition
change in proportions of heritable properties of groups of organisms over the course of generations
four forces of evolution
mutation
gene flow
genetic drift
selection
chromosomal inversion
change in chromosome structure
mutation definition
a physical change of genome (DNA)
somatic cells
NOT reproductive tissue (NOT PASSED DOWN)
germ line cells
reproductive tissue (PASSED to NEXT generation)
evolution by mutation definition
change in proportion of physical traits by the physical change of genome
evolution by mutation other important info
CANNOT cause a trait to spread
RANDOM!!!
creates raw diversity for other forces to act on
gene flow definition
transfer of genes, alleles, or traits from one population to another
organisms move from one population to another (or gametes)
change in proportion of traits in the local population
introgression
transfer of genetic material between different species of populations via hybridization followed by repeated backcrossing
genetic drift definition
any RANDOM process that affects who breeds, lives, or what genes/traits get passed down
random fluctuations in allele (or trait) frequency
locus
location on a gene
what does genetic drift do a trait among populations?
INCREASES genetic differences among populations
DECREASES genetic variation within populations
think of bell curve graph going out to the two ends
lost gene/trait
frequency of the trait/gene is 0 (left end of the two separated graphs)
caused by genetic drift
fixed trait/allele
100% of the individuals in the population have the trait/allele
caused by genetic drift
right end of the two bell curves that separated
can alleles/traits change from being lost/fixed from genetic drift?
NO they cannot change and are stuck
… can change if you evoke another force of evolution
natural selection definition
any consistent difference in fitness among phenotypically different individuals
some individuals have more babies than others because of their traits
fitness definition
number of offspring produced in a lifetime
what force of evolution leads to adaptation?
natural selection
adapt (verb)
when a population undergoes evolution by natural selection and thereby becomes better matched to the environment
individuals do NOT adapt
adaptation
an apparent match between a trait and its environment
adaptation (noun)
a trait that is matched to its environment
EVOLUTION by natural selection definition
consistent relationship for that trait, between parents and offspring, which is partially independent of common environmental effects = INHERITANCE
basic requirements
variation
consistent relationship between trait and number of offspring produced fitness differences
agent of selection definition
factor (thing) causing differences in fitness
target of selection definition
trait that is linked to fitness differences (horn length in lizard)
formula for strength of natural selection
mean (trait) of reproducing individuals - mean (trait) of entire population
Xas - Xbs
viability selection
fitness component = survival
sexual selection
fitness component = number of gametes fertilized (number of mates copulated with)
any consistent difference in # of gametes fertilized among phenotypically different traits
artificial selection
humans choose who lives and dies or who breeds based on their traits
heritability
how much of a trait, at the population level, is passed from parents to offspring
how much of the variance in a trait is due to heritable genetics
heritability graph
dots are an avg of the family
x axis = parent
y axis = offspring
slope of the line is heritability (h²)
what does the slope of the heritability graph represent?
the heritability of the trait
percentage of trait due to genetics (inherited) instead of environment
what is the breeders equation
equation that describes and predicts evolution by natural selection
quantifies evolutionary change
breeders equation (formula)
R = h² * S
what is R in the breeders equation
change in the mean of trait across generations
(how much the mean changes from parents to offspring)
evolutionary response
breeder’s equation “R” formula
x̄off - x̄bs
what does the S represent in the breeders equation
strength of selection
(change in trait value within a parental generation)
breeder’s equation S formula
x̄as - x̄bs
sexual selection definition
any consistent difference in the number of gametes fertilized among phenotypically different individuals
within sex competition
male x male competition
between sex choice
male and female choice
two parts of sexual selection
within sex competition
mate choice
Bateman gradient graph
shows number of mates on x axis and number of offspring on y axis
slope represents intensity of sexual seleciton
anisogamy hypothesis
there are differences in being male and female
females fitness constrained by investment into making offspring (why they have a less steep slope)
males will always experience stronger sexual selection
resource limitation hypothesis
differences in resource limitation dictate which sex gets a benefit from mating with more mates
the sex least constrained (less limited) experiences stronger sexual selection
speciation
evolutionary process that causes one population to become two separately evolving groups that are reproductively isolated (they can no longer interbreed)
species
group of actually potentially interbreeding natural populations that are reproductively isolated from such other groups
allopatric speciation by vicariance
populations that have been separated for long periods should be more different than species separated later and would be less likely to mate
steps of allopatric speciation
a physical barrier forms that separates what was an interbreeding population
evolution occurs differently in the two new populations. different alleles and traits get fixed in the two populations
eventually so many differences accumulate that even when given the chance the two groups can no longer interbreed and produce viable offspring
how does selection impact speciation
SPEEDS UP
can fix traits or alleles if the environment is different for the two populations (causing them to be more different)
how does mutations impact speciation
SPEEDS UP
new mutations could arise in one population but not the other
how does genetic drift impact speciation
SPEEDS UP
potentially a major force
fixes some alleles in one population and others in the other population
random, so different effects in different populations
how does gene flow impact speciation
SLOWS DOWN
mixing alleles of the two populations
prevents fixation and keeps the two populations similar
reproductive barriers
traits that have evolved while the two populations were separated that now hinder successful reproduction between the two populations
prezygotic barriers
hybrids have lower fitness than offspring from within species mating
causes populations to evolve behaviors (or other traits) that keep individuals from mating with individuals from the wrong species
prezygotic barrier examples
temporal isolation
behavioral isolation
temporal isolation
species breeds at different times (day, season, etc.)
behavioral isolation
elaborate behaviors (mating rituals) and species specific signals that prevent interbreeding between closely related populations
phylogeny
a visual representation of the evolutionary history of populations, genes, or species
reading a phylogenetic tree
node represents the last shared common ancestor
branches represent periods of stasis (no speciation)
species on the end are the species alive today
nodes can rotate and it doesn’t change the ancestors
“common descent”
darwins idea
graphing/labeling based on similar traits
hypothetically more closely related to organism that is more similar to you
parsimony
correct tree that is best supported by evidence
tree that has fewest evolutionary events
convergent evolution
when species that are not closely related evolve to have similar traits
makes creating a phylogenetic tree harder