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Natural Selection
The process by which individuals with traits better suited to their environment survive and reproduce to pass on those advantageous traits to their offspring (Charles Darwin) - Survival of the fittest
Process of Natural Selection(4)
Inheritable variation exists in a population
There is competition, or a struggle for survival - # of offspring exceeds what ENV can support
Selection - different environmental pressures exist
Individuals with the right adaptations(beneficial traits) will more likely survive, reproduce, pass on their genes.
Evolution
Occurs when allele frequency of population changes over time
Mutation
Changes in DNA sequences
Mutations, or changes in DNA sequences…(2)
Create new alleles for natural selection to act on (2)
are the only way new alleles can be generated
produce genetic variations in populations
They can be…(2)
beneficial - creates successful variation of existing trait
harmful - decrease survival or reproduction
neutral - no immediate effect
Sexual Reproduction
Reproduction involving meiosis and fertilization
Sexual Reproduction(2):
Creates new allele combinations
Increases genetic diversity without producing new alleles.
Different ways of creating diversity(3)
Crossing over
independent assortment
random fertilization
Crossing Over (2)
exchange of non-sister chromatid segments between homologous CHR during prophase I
produces recombinants, offspring that have different gene combinations than parents
Independent Assortment
Homologous CHR line up randomly and independently from one another during metaphase I.
2²³ possible gametes in humans.
Random Fertilization
Any sperm can fertilize any egg.
Overproduction of offspring(2)
Species produce more offspring that ENV. can support
ultimately, population > resources
produces competition for resources
Resources like food, space, and mates determine carrying capacity.
Allows survival of the fittest
carrying capacity
maximum population that the environment can support
Survival of the fittest
Individuals with advantageous traits (better at getting resources) are more likely to survive and reproduce
such traits will become more common over time
Selective Pressures(abiotic)
Environmental factors that affect an organism’s chances of survival and reproduction
may changes over time, altering which adaptations are beneficial for survival
abiotic factors can be(2)
positive - favor individuals with certain traits to survive and reproduce
negative - reduce the number of individuals less suited
Density Independent Factors
factors that impact a population regardless of its size
Examples of Density Independent Factors(3)
high temperatures - heatwave (coral bleaching)
low temperatures - sudden frost can kill plants, or other food sources
droughts, floods, and storms can also impact populations
Adaptations
Variations among individuals
organisms have different adaptations that allow them to be better suited to their ENV.
Intraspecific Competition and Survival
organisms with better adaptations will more likely surive in competition for the same resources
those with higher fitness will pass on their genes to the next generation.
fitness is relative, or depends on ENV
This is how natural selection changes population traits over time
Fitness
Organism’s ability to survive and reproduce
Heritability of Traits: Evolution
Change in heritable traits of a population over time, occurs via natural selection
Heritable traits
traits encoded by DNA(genes) and passed from parent to offspring
only changes in DNA like mutations can be inherited
acquired characteristics are not inherited
those gained in life (muscle, loss of limb) do not change DNA, and cannot be passed to offpsring
Environmental factors (diet, sunlight, injury) can only impact the individual not their genes.
Lamarck’s theory is incorrect
Sexual Selection(3)
Type of natural selection where certain traits increase an organism’s ability to attract a mate and reproduce.
such traits aren’t directly tied to survival
Act as signals of fitness(health, strength, and good genes)
sexual selection can produce significant differences between males and females, known as sexual dimorphism
Types of sexual selection: Intersexual Selection
Individuals of one sex (usually females) choose mates based on traits (color, dance, song)
Examples of Intersexual Selection(2)
Bird of Paradise: Males have bright plumage and complex dances that attract females, even though they may increase predation risk
Deer: Males grow large antlers, despite being energetically costly to grow and maintain
Natural Selection Experiments
Guppies are a species of fish that evolve traits as a consequence of both natural selection and sexual selection
John Endler placed guppies into artificial stream tanks with gravel of different colors.
independent variables were presence of predators(high, low, none) and color of gravel (for camouflage)
Results of Endler
Guppies with predators evolved duller colors for camouflage
favored by natural selection (survival)
guppies without predators evolved brighter colors due to female preference
favored by sexual selection (mating)
Gene Pool
Total number of alleles for all genes in a population (same species in same area, interbreeding) at a given time
Effect of gene pool size on populations
large ones increase genetic diversity, fitness, and survival
smaller ones increase chances of extinction
evolution occurs when allele frequencies in the gene pool change over time.
Allele Frequencies of Isolated Populations: Genetic Drift
Change in gene pool due to chance or random events
especially in isolated populations
larger populations maintain stability due to greater total number of alleles
Examples of Genetic Drift(2)
Bottleneck effect
Founder Effect
Bottleneck Effect
event that reduces a population size by more than 50%
natural disasters(fire, flood) or humans (overfishing)
Founder effect
small group leaves its original population to setup a new one
differs from bottlenecks, original population stays intact
Both populations will have less genetic variability
be more prone to genetic drifts
Neo-Darwinism: Allele Frequency
Prevalence of an allele, in comparison to others in the gene pool, represented as a percentage (0-1.0)
How allele frequency is evidence of evolution via natural selection
As favorable traits are passed on, the frequency of advantageous alleles increases.
alleles for less favorable traits decrease in frequency
Creation of Neo-Darwinism
Darwin proposed evolution by natural selection based on observation, unaware of genes.
modern biologists combined natural selection with genes.
Neo-Darwinism
Evolution as changes in allele frequencies caused by natural selection acting on heritable genetic variation.
Types of Selection(3)
All 3 types result in a change in allele frequency
Stabilizing Selection
Directional Selection
Disruptive Selection
Stabilizing Selection
Favors intermediate phenotype and selects over the two extremes
traits become more uniform around the average
ex. human birth weight
Directional Selection
Favors one extreme phenotype over the other
trait distribution shifts in one direction
ex. peppered moths, antibiotic resistance in bacteria
Disruptive Selection
Favors both extreme phenotypes over the immediate
trait distribution splits into 2 distinct groups
may eventually encourage speciation
ex. beak size
Conditions for Genetic Equilibrium(5) pgmms
to maintain genetic equilibrium(keep allele frequencies the same), the following five conditions must be met:
Large population size - smaller size, greater likelihood that small changes produce significant effects (genetic drift)
No gene flow - no individuals moving in/out of populations via migration
no mutations - no new alleles introduced
random mating - individuals mate without preference for traits
no natural selection - all genotypes must have equal chances or survival and reproduction
Artificial Selection
Process where humans choose individuals with desriable traits to reproduce
purpose: enhance specific traits over generations
Examples of Artificial Selection
Crop Plants - select for plants with larger fruits or pest resistance
Domesticated animals - breeding dogs for size, temperament, or coat type
Antibiotic Resistance
Bacteria have evolved resistance through natural selection, as an unintended consequence of human action
Bacteria become antibiotic resistant through mutation
When exposed to antibiotics (selective pressure), regular bacteria will die
Those with resistance will survive and reproduce
will flourish due to a lack of competition
They will further spread resistance by transferring R plasmids via bacterial conjugation
This adaptation has increased the allele frequency of the population
Lamarckism
Evolution via acquired traits
ex. trees growing asymmetrically due to wind
Sequential Evidence
Evolution is heritable, therefore change is visible in(3) DNA, RNA, AA
Sequence Comparison of Genes
fewer differences in more closely related species
Sequence differences in genes
can be accounted for by splitting ancestral species
Gene families like Hox Gene
Hox Gene
Determines body plan during development
found in all ORG with a clear head-to-tail axis
Selective Breeding