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Speciation
the process by which species are created and destroyed
What are the two ways things go extinct?
cease to exist or go bye bye
organism goes into an evolutionary trajectory and evolve into something else, so their previous form does not exist anymore
Primitive
implies ancestral and old when compared to geologic time (plesiomorphic traits)
Derived
implies recent and advanced (Synapomorphic traits)
Speciation Process
everything that was primitive can evolve and become derived
3 methods of speciation theories
allopatric speciation
sympatric speciation
parapatric speciation
Allopatric Speciation (different land)
a population of a species becomes isolated into 2 sub-populations, organisms then evolve to the specificities of the environment to which they have become isolated
environments are not the same, they are different, animals have to start to respond to the difference
Clinal Variation
variation across the gradient where there are two extremes on the extremes and in the middle there is an average, anything with a large distribution
Allopatric speciation isolation events
Vicariant Speciation
Founder event Speciation
Vicariant Speciation mechanism
climatic or geologic change results in a population becoming split
ex: mountain ranges, rivers, glaciers, watershed shifts, and climate change
Sympatric Speciation (Same Land)
speciation without geographic isolation can occur
occurs on the same land, where generalists create founding stock for future speciation events
speciation follows an evolutionary trend behaviorally towards specialization
Progenitor Form
primarily found in sympatric
Parapatric Speciation
speciation with both geographic and behavioral modalities of speciation
geographically, species are isolated
though they can make contact along a border line, neither species crosses
Adaptive Radiation
adaptations radiate out into new phenotypes and genotypes when an ancestral taxa occupies several new niches or habitats and results in a net gain in species
Microclimate
a very small area that has a different climate
Darwin’s proposal on how long it takes species to form
proposed speciation was a long, gradual process taking thousands to millions of years to occur
Gradualism
the prevailing theory of casual evolution
Gradualism types
Populational Gradualism: new traits become established in a population by increasing their frequency initially from a small fraction of the population to the majority
Phenotypic Gradualism: new traits, even those that are strikingly different from ancestral ones, are produced in a series of small incremental steps
Morphological Divergence
physical representation of gradualism as a graph
What does gradualism NOT explain?
how taxal groups with long evolutionary legacies form or how taxal groups which rapidly form and appear in the fossil record
2 strikes against gradualism
mass extinctions
gaps in the fossil record
Punctuated Equilibrium
relatively recent, theorizes that speciation can occur rapidly, especially ecosystems that have experienced a biological crisis (extinctions)
What happens if several niches are suddenly opened?
speciation rates across taxa increase to occupy those niches
in response to this, speciation occurs rapidly (this can explain gaps in the fossil record and can explain raised levels of diversity immediately following mass extinctions
Natural Selection
“only the strong survive, the weak perish”
strong = ability to reproduce and get to the next generation
From what perspective do biologists view natural selection:
2 faces working towards one goal:
one face points towards an organism’s genotype
from the genotype, the other face is displayed in an organism’s phenotype
Genes that are advantageous
increase an organism’s fitness (which is the goal)
Artificial Selection
to better understand selection in nature, biologists have experimented with selection in controlled settings
How do you complete a cycle of natural selection?
the average fitness of the offspring needs to change
Species phenotypes
species do not have an ideal phenotype, because time does not stop
What occurs when selection is weak or variable in direction?
it cannot act effectively on a population
What is the additional factor that limits natural selection’s impact?
the number of deaths occurring to sexually mature individuals in a population
each individual must ensure that it at least replaces itself to maintain current population levels
What may an organism’s phenotype might be selected for
growth under normal conditions
growth with limited nutrition
growth with limited water intake
disease resistance
What occurs at different times over the course of the species’ existence?
different phenotypes will be successful
What is the species long-term resistance dependent on?
the possibility of the presence of all 4 phenotypes