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What is symbiosis?
Close physical association between two or more different organisms
Three types of symbiosis?
Mutualism → both benefit
Commensalism → one benefits, other unaffected
Parasitism → one benefits, one harmed
Why is symbiosis important evolutionarily?
Responsible for major innovations in life (endosymbiosis)
What do plants gain?
Nutrients (N, P)
Protection from pathogens
Pollination & dispersal
Most limiting nutrients?
Nitrogen (N) and Phosphorus (P)
Who can fix nitrogen?
Prokaryotes ONLY
Reaction of nitrogen fixation?
N₂ → NH₄⁺ (usable form)
What bacteria are involved? symbiosis legumes
Rhizobium
Where does fixation occur? symbiosis legumes
Root nodules
Why is nitrogen fixation tricky?
Requires low oxygen (anaerobic conditions)
How is nitrogen fixation maintained?
Plant hemoglobins regulate O₂
What are mycorrhizae?
Symbiosis between fungi + plant roots
Plant gets:
↑ nutrient uptake (especially phosphorus)
Protection
Larger effective root system
Fungus gets:
Sugars (carbon)
Types of Mycorrhizae
Endomycorrhizae (VAM)
Ectomycorrhizae
Endomycorrhizae (VAM)
Fungus enters root cells
Most common (~65%)
Exchanges happen via arbuscules
Ectomycorrhizae
Fungus surrounds root
Does NOT enter cells
Common in woody plants
Do fungi cross the endodermis?
No
What are fungal endophytes?
Fungi living inside plant tissues
Function fungal endophytes?
Protect against herbivores
Increase stress tolerance
Pollination Symbiosis
Fig + wasp
Yucca + moth
Why is agriculture symbiosis?
Humans + crops both benefit
Humans → food
Plants → protection, spread
Why do plants perceive environment?
To regulate:
Germination
Flowering
Growth
Dormancy
four categories of environemntal perception
Tropisms
Photomorphogenesis
Circadian rhythms
Photoperiodism
What are tropisms?
Growth toward/away from stimulus
Types of tropisms
Phototropism → light
Gravitropism → gravity
Thigmotropism → touch
What detects light?
Phototropins (blue light receptors)
What happens in phototropism mechanism?
Auxin redistributes → uneven growth → bending
Auxin Rule
Shoots: ↑ auxin → ↑ growth
Roots: ↑ auxin → ↓ growth
What senses gravity?
Statoliths (amyloplasts)
Mechanism of gravitropism
Settle → signal → auxin redistribution → bending
Direction of Growth
Roots → positive gravitropism (down)
Shoots → negative gravitropism (up)
What triggers Thigmotropism?
Touch → Ca²⁺ signaling → auxin redistribution
what is photomorphogenesis
Development controlled by light
Example: Etiolated (dark-grown) vs light-grown plants
Types of Photoreceptors
Phototropins → blue light
Phytochromes → red/far-red
Cryptochromes → blue/UV
Two forms of phytochrome system?
Pr (red-absorbing)
Pfr (far-red absorbing, active)
what triggers shade avoidance?
Low red:far-red ratio (shade)
Effects:
Taller growth
Less branching
Earlier flowering
What are circadian rhythms?
24-hour internal biological cycles
What is photoperiodism?
Response to day length
Key molecule for flowering control?
Florigen (FT protein)
What gene regulates flowering?
CONSTANS (CO) → activates FT