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Altruism
an action that reduces an individuals own fitness while increasing the fitness of another
inclusive fitness
direct fitness (offspring) and equivalents gained through support (kin)
Nash equilibrium
-the best response given another's response
-always play the selfish card
-unstable
evolutionarily Stable strategy
-a strategy that cannot be invaded by a rare mutant using an alternative strategy
-special case of Nash equilibrium
cooperatively breeding
-some members forego reproduction
-devote energies toward rearing of young of others
eusoical
-individuals cooperate in the care of young
-reproductive division of labor
2 hypotheses of why altruism happens
1. selection at the level of the family (kin selection)
2. reciprocity
kin selection
-acting on individuals related through recent, common descent
-depends of relatedness (percentages of shared allels)
hamilton's rule
fitness=w-c+rb
act altruistically if c
reciprocal altruism
altruism is rewarded with altruism, defection punished by defection
cost v benefit of helping
cost= fitness when not helping - fitness when helping
benefit= fitness when helped- fitness when not helped
genera
earliest classification system which grouped animals and plants into units
taxon
organisms at a particular level in a classification system
scientific names
first part is genus, second is specific epithet (species)
Hierarchical Classification
Domain, Kingdom, Phylum, Class, Order, Family, Genus, Species
John Ray
first definition of a species
all the individuals that belong to it can breed with one another and produce fertile offspring
biological species concept
-defines species as groups that are reproductively isolated
-doesn't work for asexual reproductive organisms
phylogeny
the evolutionary history of an organism and its relationship to other species
cladistics
phylogeny according to similarities derived from a common ancestor
clade
a group of organisms that share a derived character
6 kingdoms
Animalia, Plantae, Fungi, Protista, Archea, Bacteria
3 domains
Archaea, Bacteria, Eukarya
which prokaryotic kingdom is closer to eukaryotes
archaea
Archaean characteristics
-unique cell walls, lipids, and rRNA sequences
-some genes have introns
-found in extreme environments
When did eukaryotes appear
1.5 billion years ago
endosymbiosis
bacteria entering an early eukaryotic cell created a mitochondria and chloroplasts
when did prokaryotes first appear
3.7 billion years ago
evolutionary tree
a diagram that reflects evolutionary relationships among groups of organisms based on similarities in DNA, physical features, biochemical characteristics, or a combo
node
marks when an ancestral group split into two separate lineages
fungi includes
yeasts, molds, mushrooms
who were the first terrestrial colonists
algae-like plants
waxy cuticle purpose
retain moisture
gymnosperms
-first plants to evolve pollen and seeds
-removed dependence on water for fertilization
angiosperms
-first plants to evolve flowers
-most dominant and diverse group of plants
order of main events
1. 650 mya life in water
2. 540 mya Cambrian explosion
3. 480 mya algae-like plants make it to land
4. 460 mya fungi make it to land
5. plants on land
6. 360 mya end of the Devonian period
7. 400 mya animals on land
8. 365 mya amphibians on land
9. 230 mya reptiles most common
10. dinosaur age
how many mass extinctions have there been?
5
causes of mass extinction (4)
climate change, massive volcanic eruptions, changes in composition of marine and atmospheric gases, and sea level changes
Cretaceous extinction event
-65 mya
-wiped out 3/4 of plant and animal species (including non-avian dinos)
-massive comet/asteroid in Gulf of Mexico
adaptive radiation
-a group of organisms expand to take on new ecological roles and form new species and higher taxonomic groups
-happen after mass extinctions
example of adaptive radiation
mammals diversifying after extinction of dinosaurs
ratio of human cells to microbial cells
1:1
affects of the human microbiome
-gut health
-brains
-body odor
-risk of infection
-attractiveness (to people and mosquitos)
who was the first to branch off from the shared ancestor
bacteria
prokaryotes
-single-celled organisms
-single loop of DNA
-able to reproduce rapidly
prokaryote basic structural plan
-protective cell wall
-some have a capsule around the cell wall, and/or pili, and/or a flagella
what do capsules do
help bacteria to evade the immune system
what do pili do
-help organisms link together and attach to surfaces
what does a flagella do
help propel bacteria through liquid
thermophiles and halophiles
- archaeans
-heat and salt lovers
quorum sensing
-a system of cell-to-cell communication
-allows them to sense and respond to other bacteria in accordance with the density of the pop.
binary fission
-method in which prokaryotes reproduce
-asexual reproduction
-splitting of DNA
prokaryotic forms of reproduction
binary fission, horizontal gene transfer, bacterial conjugation, sporulation
horizontal gene transfer
-indirect transfer of genes by capturing bits from the environment and incorporating them into their own genes
bacterial conjugation
-direct transfer of genes between bacteria
sporulation
-formation of thick-walled dormant structures (spores)
-can survive harsh temps
-allow microbes to survive until conditions are favorable to reproduce
metabolic types
-aerobes (need oxygen)
-anaerobes (some are methanogens, some perform fermentation)
feeding types
chemoheterotrophs, photoheterotrophs, chemoautotrophs, photoautotrophs
nitrogen fixation
Process of converting nitrogen gas into ammonia (bacteria)
dysbiosis
when the human microbiome shifts out of balance
features of eukarya
-true nucleus
-DNA enclosed in 2 concentric layers of membranes
-organelles
Protists
-defined by exclusions
-most single-celled eukaryotes
-protozoans and algae
-flagella, cilia, pseudopodia
-many are heterotrophs
protozoans
nonphotosynthetic and motile protists
algae
photosynthetic and may/may not be motile
how to motile protists move?
-swim by flagella or cilia
-crawl by pseudopodia (false feet)
why can multicellular organisms grow large?
they can gather resources more effectively which allows for production of more surviving offspring
plants
-evolved from green algae
-multicellular autotrophs
-vascular tissue
bryophytes
-liverworts and mosses
-ancestors of first plants
-simple plants, small
-absorbed water through a wicking action
vascular tissue
a network of tubelike structures specialized for transporting fluids
-first plants with these were ancestors of ferns
ferns
-vascular tissue
-lignin
seeds
-consist of the plant embryo and stored food supply
-came from gymnosperms
fungi
-absorptive heterotrophs
-zygomycetes, ascomycetes, basidiomycetes
-protective cell wall, store food energy in the form of glycogen
zygomycetes
mold species
ascomycetes
-diverse group
-sac fungi
basidiomycetes
club fungi
truffles
-fruiting body of a type of ascomycetes
yeasts
-single-celled fungi
-produce alcohol and carbon dioxide
fungi structure
-body is a network of thin filaments
-provides large surface area to absorb nutrients
-penetrates food source
-secrete enzymes
importance of fungi
-economically (bread, wine, beer)
-gastronomically (edible mushrooms)
-medically (penicillin)
hyphae
branching fungal threads that absorb nutrients from the environment
mycelium
mass of hyphae forming the body of a fungus
fungal reproduction
both asexual and sexual
asexually through fragmentation
mycorrhizal fungi
fungi that surround plant roots and help plants obtain water and minerals
lichen
-mutualistic association between a fungus and a photosynthetic microbe
-microbe provides sugar and carbon compounds
-fungi provides lichen acid for protection
Neanderthals
-closest human relative
-humans contain some of their DNA
-humans and them interbred
-same genus as humans
animals
-multicellular ingestive heterotrophs
-first evovled 700 mya
sponges
-asymmetrical body plan
-most ancient animal lineage
body symmetry
-all animals except sponges
-radial (vertical plans across center)
-bilateral (one plane vertically in two halves)
chordates
-dorsal notochord
-dorsal nerve cord
dorsal notochord
-a flexible rod along center of the body crucial for development
-evolved into discs between vertebrae
vertebrates
all chordates with backbones
evolution of vertebrates
1. jawless fish (cartilage skeleton)
2. hinged jaws
3. replacement of cartilage to bone
Class Mammalia
-body hair, sweat glands, mammary glands
-eutherians, marsupials, monotremes
eitherians
-offspring are nourished inside the mother's body via placenta
-born in a relatively well-developed state
marsupials
-simple placenta
-offspring are born early and complete development in an external pocket
monotremes
-lay eggs
-no placenta
-platypus and several echidna
primates
-flexible shoulder and elbow joints
-5 fingers and toes
-opposable thumbs
-flat nails (not claws)
-large brains in relation to body size
hominids
-the ape family
-use of tools, capacity for symbolic language, deliberate acts of deception
-includes humans
hominins
-human branch of ape family
-one or more humanlike feature (thick tooth enamel, upright posture)
-includes neanderthals
what distinguishes hominins from hominids
being bipedal
mitochondrial-DNA inheritance
-occurs only from egg (not sperm)
-passed down virtually unchanged
-can be tracked from 1 generation to another and 1 species to another