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what extra features did vertebrates evolve?
bony living skeleton
made of secreted collagen fibres + calcium phosphate crystals
osteoblasts make bone, osteoclasts break down bone, and osteocytes are strain/crack sensors
vertebra are formed from fusing of bone produced at the sclerotome, the inner part of the somites (dorsal, segmented mesoderm- unlike the ventral, lateral plate mesoderm)
neural crest cells
individual cells migrating from the inner section of the infolding of the dorsal hollow nerve cord
these form: sensory nerves to the skin, pigment cells, nerves in the gut for peristalsis, autonomic nervous system, some blood vessel muscle, and the bone + cartilage of the skull
red blood cells
invertebrates only have a few cells in the blood for phagocytosis, but vertebrates have red cells with haemoglobin
adaptive immune system
the innate immune system of invertebrates recognises common pathogens, but the adaptive immune system in vertebrates changes with the environment by dna rearrangement
this is due to antibody genes and t-cell receptors
these traits allowed early vertebrates to evolve into big, active, swimming, hunting predators

what features of vertebrates became more complex?
sense organs of the head
olfactory- chemical sensing
optic- light sensing
otic- mechanical sensing
larger brain
forebrain- olfaction + thinking in some vertebrates
midbrain- optic lobes
hindbrain- neural crest cells in this region make the cartilagenous gill skeleton:
pharyngeal slits
supports gills in fish
the first slit is the jaw bone, the second is a jaw support bone
these traits allowed early vertebrates to evolve into big, active, swimming, hunting predators

describe the phylogeny of the vertebrates
vertebrates are a subphylum of the chordate phylum in the deuterostomia clade of bilaterian animals
the first lineage to evolve was the agnatha (lamprey + hagfish), before the gnathostomata developed jaws + paired fins:
chondrichthyans- cartilagenous fish eg. sharks + rays
osteichthyes- bony fish
actinopterygians- ray-finned fish eg. most fish
sarcopterygians- lobe-finned fish + tetrapods (terrestrial)
the ‘fish’ is a paraphyletic group/grade, not a clade- it contains the most recent common ancestor but not all the descendants (no tetrapods)

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