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Population Density
N per area/volume
Population Dispersion
indiv spacing pattern within pop boundaries
Mark-Recapture Method
sampling technique to est pop size; N = sn/x
Dispersion Types
clumped, uniform, random
Clumped Dispersion
indivs in patches; resource availability/behavior (most common)
Uniform Dispersion
evenly spaced indivs; antagonistic interactions/territoriality
Random Dispersion
unpredictable spacing; w/o strong attractions/repulsions
Demography
study of pop vital statistics (birth/death/migration) & changes over time
Life Table
summarizes survival & reproductive rates of specific age-groups (cohorts)
Type I Survivorship Curve
low early/mid death rate, > late death rate
Type II Survivorship Curve
constant death rate over lifespan
Type III Survivorship Curve
>/< early/late death rate
Exponential Population Growth
pop growth under ideal/unlimited env -> J-shaped curve
Intrinsic Rate of Increase (r)
per capita rate of pop growth at each instant in time
Exponential Growth Equation
dN/dt = rN
Carrying Capacity (K)
max env sustainable pop #
Logistic Population Growth
per capita growth rate -> 0 as pop nears K -> S-shaped curve
Logistic Growth Equation
dN/dt = rN((K - N)/K)
Life History
traits affecting reproduction/survival schedule (when/how often/how many)
Semelparity
1-time "big-bang" reproduction -> death
Iteroparity
rep repro over lifespan
Reproductive Trade-offs
lim resources -> compromise btwn offspring # vs provisioning/parental care
K-Selection
selection for traits advantageous at high pop density (near K)
r-Selection
selection for traits maximizing reproductive success at low pop density (un-crowded)
Density-Independent
birth/death rate !change w/ pop density
Density-Dependent
birth/death rate changes w/ pop density -> - feedback
Density-Dependent Regulation Mechanisms
resource competition, territoriality, disease, predation, toxic wastes, intrinsic (hormonal) factors
Population Dynamics
biotic/abiotic interactions -> pop # fluctuations ot
Population Cycles
regular boom-and-bust fluctuations
Metapopulation
local pops linked by immigration/emigration
Demographic Transition
shift from high birth/death rates to low birth/death rates
Age Structure
relative # of indivs of each age in a pop -> future growth trends
Ecological Footprint
aggregate land/water area req to produce resources consumed & absorb waste
Density Estimation Methods
sample plots, indicators (nests/burrows/tracks/feces), mark-recapture
Molecular Demographics
PCR/genetic profiles -> track individual reproduction w/o disturbance (e.g., loggerhead turtles)
Stair-Stepped Survivorship
brief periods of > mortality followed by < mortality
Population Size Change Equation
dN/dt = Births + Immigrants - Deaths - Emigrants
Logistic Model Caveat
assumes instantaneous adjustment to K; reality -> time delay -> temporary K overshoot
Metapopulation Dynamics
balance of local patch extinctions & immigrant recolonizations
Metapopulation Genetics
heterozygote advantage -> > flight -> > new patch colonization
Zero Population Growth Configurations
high birth rate - high death rate / low birth rate - low death rate
Infant Mortality
infant deaths per 1,000 live births; high rate -> parents + reproductive rate to compensate
Life Expectancy at Birth
predicted avg lifespan; recent < driven by disease (AIDS) & social upheaval
Demography Focus
often ignores males -> tracks only females giving rise to new females
Mark-Recapture Assumption
marked/unmarked individuals have = probability of capture; marked fully mix back in; no births/deaths/migration during interval
Conservation Logistic Model
used to estimate critical pop size below which endangered species may become extinct
Offspring Trade-off (Plants)
many small seeds for far wind dispersal / few large seeds w/ > endosperm nutrients for establishment
Density-Dependent Predation
> prey pop -> < hiding spaces -> + predation death rate
Density-Dependent Intrinsic Factors
high density -> aggressive interactions/hormones -> delayed sexual maturation & - immune system even if food is abundant
Human Pop Growth Rate
peaked at 2.2% in 1962; < w/ China famine; currently <
Age Structure Consequences
bottom-heavy -> rapid growth & future employment/education problems (e.g., Zambia) / small-base -> fewer working-age people to support retired "boomers"
Global Hectare (gha)
unit of biologically productive area; sustainable limit = 1.7 gha/person, but US avg = 8 gha/person ->unsustainable
Interspecific Interactions
competition, exploitation, positive interactions
Competition (-/-)
diff species individuals use same limiting resources -> < survival/reproduction
Competitive Exclusion
1 species uses resources > efficiently -> local elimination of inferior competitor
Ecological Niche
specific set of biotic & abiotic resources an organism uses
Resource Partitioning
niche differentiation -> enables ecologically similar species to coexist
Fundamental vs. Realized Niche
potential niche vs portion actually occupied (due to competition)
Character Displacement
sympatric pops diverge > in morphological traits than allopatric pops
Exploitation (+/-)
1 species benefits by feeding on/harming other (predation, herbivory, parasitism)
Aposematic Coloration
warning coloration (e.g., poison dart frog) -> + predator avoidance
Cryptic Coloration
camouflage -> makes prey difficult to see
Batesian Mimicry
harmless species mimics harmful species
Müllerian Mimicry
2+ unpalatable species mimic each other
Herbivory (+/-)
herbivore eats plant/alga parts -> harms !kills
Parasitism (+/-)
parasite derives nourishment from host -> host harmed
Endoparasites / Ectoparasites
live within host body / feed on external host surface
Positive Interactions (+/+ or +/0)
1+ species benefits & neither harmed
Mutualism (+/+)
interaction benefits both species
Commensalism (+/0)
interaction benefits 1 species & !harms/helps other
Species Diversity
variety of diff organisms; species richness + relative abundance
Species Richness
# of diff species in community
Relative Abundance
proportion each species represents of all individuals
Diversity Benefits
> productivity/biomass, > stability, > resistance to introduced species
Biomass
total mass of all organisms in habitat
Food Web
linked food chains diagramming trophic relationships
Energetic Hypothesis
food chain length limited bc energy transfer inefficiency (~10% transferred)
Foundation Species
highly abundant/large species w/ strong community effects (e.g., trees)
Keystone Species
!abundant but exert strong control via pivotal ecological roles (e.g., sea stars)
Ecosystem Engineers
species that dramatically alter physical env (e.g., beavers)
Bottom-Up Control
nutrient supply/lower trophic levels limit higher trophic level abundance
Top-Down Control
higher trophic level consumers limit abundance of lower trophic levels (trophic cascade)
Nonequilibrium Model
communities constantly changing post-disturbance
Disturbance
event (storm/fire/human) removing organisms or altering resource availability
Intermediate Disturbance Hypothesis
moderate disturbance levels -> > species diversity than low/high levels
Ecological Succession
predictable species replacement sequence post-disturbance
Primary Succession
begins in lifeless area w/o soil (e.g., glacial moraine)
Secondary Succession
recolonization post-disturbance where soil remains (e.g., post-forest fire)
Succession Processes
early species can facilitate, inhibit, or tolerate later arriving species
Latitudinal Gradients
tropics have > species diversity due to older evolutionary history & > climate/evapotranspiration
Species-Area Curve
> geographic area -> > species # (more diverse habitats/microhabitats)
Island Equilibrium Model
species # balance between new immigration & existing extinction
Island Equilibrium Factors
immigration < & extinction > as species # +; island size & distance from mainland determine rates
Pathogen Impact
highly virulent in new habitats bc new hosts lack evolutionary resistance
Zoonotic Pathogens
diseases transferred from animals to humans (3/4 of emerging human diseases)
Vector
intermediate species (e.g., ticks/mosquitoes) transferring zoonotic pathogen to humans
Plant Herbivory Defenses
physical (thorns/spines) & chemical (strychnine/nicotine/tannins)
Parasite Behavior Modification
parasite alters host behavior to > chance of reaching next host (e.g., infected crustacean leaves cover)
Shannon Diversity Index (H)
H = -(pA ln pA + pB ln pB + …); higher H = > community diversity
Molecular Diversity Tools
PCR amplifies ribosomal DNA -> gel electrophoresis -> generates DNA profile for invisible microbes