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evolution
heritable change in characteristics of a population or species between generations
microevolution
changes in single gene in a population over time
macroevolution
formation of new groups of species
species
group of related organisms that share a distinct form, sexually reproducing species interbreed
population
members of the same species likely to encounter each other and will interbreed
empirical thought
relies on an observation to form an idea/hypothesis, a big shift towards empirical thought happened in the 1600s
john ray
the first to carry out a thorough study of the natural world, developed an early classification system used in modern species concept, mid to late 1600s, neither of his ideas proposed evolution
1700s scientists found lifeforms arent
fixed
george buffon
“life forms change over time”
jean baptiste lamark
found that some animals remain the same while others change, believed living things evolved towards human perfection
uniformitarianism hypothesis
slow geological processes lead to substantial change, earth is older than 6k years (against ideas at time)
thomas malthus
economist, said only a fraction of any population survives and reproduces
charles darwin
noticed distinctive traits of island species that allowed them to better exploit their environment
charles darwin accomplishments
formulated theory of evolution mid 1840s, spent years studying barnacles, worked with alfred wallace to publish joint similar papers, published them titled on the origin of species, and was the crews naturalist on the HMS beagle voyage
Descent with modification
evolution based on variation within a given species with heritable traits, genetic basis was not yet known
natural selection
more offspring produced than can survive, competition for limited resources, better traits flourish and reproduce
grants
observed natural selection in finches, focused much work on island daphne major, a moderately isolated undistrurbed habitat and its resident finches
evidence of evolutionary change
fossil record, biogeography, convergent evolution, selective breeding, homologies including anatomical, developmental, and molecular
Fishapod
illuminates steps leading to evolution of tetropods
transitional form
provides link between late and early forms
fossil record spans how long
50 million years
terrestrial tetropod evolved to aquatic animals lacking hind limbs
order: whales→ dolphins→porpoises
biogeography
study of the geological distribution of extinct and modern species
isolated islands and continents have
distinct plant and animal communities
endemic
naturally found only in a particular location
evolution of major animal groups corrolate with
known changes in distribution of landmasses on earth
first mammals arose
225 million years ago when australia was still connected
why does australia have no large terrestrial placental mammals
has marsupials, most of which are exclusive to australia because pangea led australia to be isolated and placental mammals never outcompeted the marsupials
plate tectonics theory
the crust divided into giant, irregularly shaped plates that float on top of the mantle (includes both ocean and land)
continental drift
movements in the mantle that cause the plates to move
boundaries of some plates are hotspots for
geological activity (earthquakeks etc)
earthwuakes signal
plates are scraping past or colliding with each other
continental drift majorly impact evolution by
reshaping the physical features of the planet and altering the environment organisms live in
when was pangea formed
250 million years ago, reduced shoreline, deepened ocean basins, lowered sea levels
convergent evolution
two different species from different lineages show similar characteristics due to similar environments and arent related evolutionarily
selective breeding
programs and procedures designed to modify traits in domesticated species (artificial selection)
homology
descent from a common ancestor, can be anatomical, developmental, molecular
anatomical homology
ex: multiple species sharing phalanges, derived from a common ancestor
vestigal structures
anatomical with no apparent function but resembles ancestors (tailbone)
developmental homology
similarities in embryotic stages, presence of gill ridges in humans suggests evolution from fishy ancestor, human embryos have long bony tails
molecular homology
similarities in cells on a molecular level shows common ancestor, all living species use dna, certain biochemical pathways found in nearly all species
sequences of closely related species tend to be more ___ than distantly related species
similar
homologous genes
two genes derived from the same ancestral gene
orthologs
genes in different species that evolved from a common ancestral gene by speciation, typically retaining the same function, reveals molecular details of evolutionary change
two sequences can be ____ but not ___ because of accumulation of random mutations
similar, identical
gene duplicates
produce gene families
gene families are
two or more paralogs within the genome of a single organism (ex, globin genes, allow for specialized function, expression at different times or in different tissues).
ortholog genes are related by
vertical descent from a common ancestor, encode proteins with the same function in different species
paralogs
homologous genes that have evolved by duplication and code for a protein with similar but not identical functions
horizontal gene transfer
exchange of gene material among different species (common)
where does horizontal gene transfer take place
prokaryotic to eukaryotic, vice versa, between prokaryotes, between eukaryotes, and is widespread among bacteria
evolution also occurs at the ____ level
genomic
evolution at the genomic level
involves changes in chromosome structure and number (humans have one large chromosome while apes have it divided into separate chromosomes)
chromosome 3 between humans and orangutans are very similar but orangutans have a large _____ of the chromosome which may have established orangutans as a new species
inversion
population genetics
studies genes in populations and genotypes, aims to understand extent of genetic variation, why how, and what changes happen
also helps understand how genetic variation is related to phenotypic variation
population
group of individuals of the same species that share environment and can interbreed
discrete populations
population that occupies wide geographical range is divided into smaller populations
changes in population size and location =
changes in genetic composition
genes are usually _________
polyphormic
polyphormic genes are
two or more variations for a given character, due to two or more alleles that influence phenotype
polyphormic gene definition short
2 or more alleles
monophormic gene
predominantly single allele
single nucleotide polymorphisms (SNPs)
smallest type of genetic change in a gene, most common type of change, responsible for 99% of variation in human gene sequence
large healthy populations exhibit high level of genetic _____
diversity
what are analysis of SNPs important for
personalized medicine
allele frequency equation
number of copies of a specific allele in a poopulation / total number of all alleles for that gene in the population
genotype frequency equation
number of individuals with a particular genotype in the population / total number of individuals in population
hardy weinburg equation
p²+2pq+q²=1
p²
genotype frequency of C^dominantC^dominant homozygotes
q²
genotype frequency of C^recessiveC^recessive homozygotes
2pq
2 times the genotype frequency of C^rC^d heterozygotes
hardy weinburg equillibrium
predicts that allele and genotype frequencies will remain the same provided a population is not evolving through generations
what standards must a population be in to fit the hardy weinburg equllibrium?
population must not be affected by evolitionary mechanisms that change allele and genotype frequencies including:
no mutations must occur
no natural selection
population is too large for alleles to change due to random sampling error
no migration occurs between different population
random mating
does hardy weinburg equllibrium ever happen
no, but its still useful for approximating some genes
microevolution
changes in a populations gene pool generation to generation
why does microevolution occur
a new genetric variation is introduced: mutations, duplications, horizontal transfer
is microevolution a major factor in allele frequencies
no (microevol)
what evolutionary mechanisms alter the prevalence of an allele of genotype
natural selection, non random mating (increases potential for widespread genetic change)
natural selection
process in which beneficial traits that are heritable become more common in successive generations, over time resulting in adaptations that promote survival and reproduction in a particular environment
fitness
relative likliness the genotype will contribute to the next generation
reproductive success
likelihood of an individual contributing fertile offspring to the next generation, attributed to two categories of traits
2 categories of traits for reproductive success
1) traits directly associated with reproduction, 2) characteristics that make organisms more likely to survive to reproductive age
modern description of natural selection
within a population, allelic variation arises from random mutations that cause different dna sequences, some alleles encode proteins that enhance survival or reproductive capability compared to other members of the population (these individuals are more likely to survive and reproduce for their alleles to be added to next gen gene pool)
over many generations, allele frequencies may significantly alter _____ of a population
characteristics
mean fitness of a population
average reproductive success of a population
as individuals with higher fitness values become more prevalent,
natural selection increases the mean fitness of the population
sexual selection
directed at traits of sexually reproducing species that make it more likely for individuals to find or choose a mate or engage in successful mating, in many species affects male population more than female
sexual dimorphism
significant different between the morphologies of the two sexes within a species
directional selection
individuals at one extreme of a phenotypic range have greater reproductive success in a particular environment
initiators of directional selection
new alleles of higher fitness are introduced with prolonged environment change
stabilizing selection
favors the survival of indiciduals with intermediate phenotypes, extreme values of a trait are selected against: ex, too big or too small of a litter decreases survivability and productivity, so an intermediate is selected for
diversifying selection
favors the survival of two or more different genotypes that produce different phenotypes, likely to occur in populations that occupy heterogenous environments. members of the populations can freely interbreed
balancing selection
maintaints genetic diversity
balanced polymorphism:
two or more alleles are kept in balance, and therefore are maintained ina population over the course of many generations.
intrasexual selection
males directly compete for mating opportunities or territories, between members of same sex, horns and such in males as a result
intersexual selection
members of one sex, usually females, choose their mates from individuals of the other sex based on desirable characteristics, often results in showy characteristics for males
cryptic female choice
genetic tract/egg selects against genetically related sperm to inhibit inbreeding
what does sexual selection explain
traits that decrease survival but increase reproductive success ex colored fish
results of sexual selection
in places where preadators are few, there are more colorful males and opposite for high amounts of predators, relative abundance of brightly and dully colored males depends on the balance between sexual selection
cichlidae have over ____ species
3000