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natural selection was developed by _____________.
charles darwin
natural selection is established due to __________ in the population and competition for ____________
variation, resources
What is evolutionary fitness?
Reproductive success (ability to pass on genes)
organisms with more favorable traits are more likely to survive and produce more offspring to pass on their traits to the next generation
What happens to organisms with favorable traits?
More likely to survive and reproduce → traits increase in population
Give two examples of natural selection
Peppered moths, antibiotic resistance
What is a Lamarckian idea (and why is it wrong)?
Traits acquired during life are inherited (incorrect—traits must be genetic)
What is disruptive selection?
Favors extreme phenotypes, selects against intermediate
What is stabilizing selection?
Favors intermediate phenotype, selects against extremes
What is directional selection?
Favors one extreme phenotype
What is artificial selection?
Organisms with certain traits are bred until population has that trait. Humans breed organisms for specific traits. Humans affect variation in the population.
Examples of artificial selection
Dog breeds, corn from maize, wild mustard → broccoli/cabbage/etc.
Natural selection can only act on..
existing features
What creates new genetic variation?
Mutations
Migration
gene flow (addition or removal of alleles from a population)
What is genetic drift?
Nonselective process in small population changes allele frequency
Random change in allele frequencies
Ex) bottleneck and founder effect
Mechanisms of Genetic Variation / Drivers of Evolution
Mutations – new genetic variation
• Migration – gene flow (addition/removal of alleles from population)
• Genetic drift – nonselective process in small populations changes allele frequencies
• Bottleneck effect – population size reduced to small number of individuals
• Founder effect – population separated from other members of population
Types of genetic drift are
bottleneck effect and founder effect
What is the bottleneck effect?
Population drastically reduced → less genetic diversity
What is the founder effect?
Small group starts new population → different allele frequencies
population separated from other members of population
Why is genetic diversity important?
Increases survival under environmental change
What happens to populations with low diversity?
Higher risk of extinction
Can an allele be both beneficial and harmful?
Yes, depends on environment
Alleles that are adaptive in one environmental condition may be _________ in another because of different ________ __________.
deleterious, selective pressures
What is Hardy-Weinberg equilibrium?
a principle stating that allele and genotype frequencies in a population will remain constant from generation to generation in the absence of other evolutionary influences. It serves as a null model for non-evolving populations, assuming random mating, no mutation, no migration, no selection, and a large population size.
Five conditions (Five Fingers of Evolution)
Large population
Random mating
No mutations
No gene flow (migration / emigration)
No natural selection
The Hardy-Weinberg equilibrium is never met but serves as a ____ ______________ for non-evolving populations, assuming random mating, no mutation, no migration, no selection, and a large population size.
null hypothesis
Equation for allele frequency
p + q = 1
Equation for genotype frequency
p² + 2pq + q² = 1
Any change in allele frequencies between generations is called
microevolution
Evolution (is / is not) occurring if p and q change!
is
What is 2pq?
Heterozygous genotype frequency
Hardy-weinberg is a valuable tool for predicting frequencies if allele frequencies (are/are not) changing! This means it will be correct if the 5 criteria are met.
are not
Null hypothesis
hypothesis which the researcher tries to disprove. Assumption that there is not relationship between the independant and dependant variable in a study.
The conditions of HWE are never met, but in experimental studies, they provide a null hypothesis for determining which factors are changing allele frequencies.
what is gene flow
movement of alleles between populations
p²
proportion of individuals that show the homozygous dominant phenotype (AA)
2pq
proportion of individuals with heterozygous genotype (Aa)
q²
proportion of individuals with the homozygous recessive genotype
How are fossils dated?
1) the age of the rocks where a fossil is found
2) the rate of decay of isotopes (incl.carbon-14)
3) geographical data.
What is biochemical evidence for evolution?
DNA or protein
Comparison of the number of differences
What is biogeography?
distribution of species and ecosystems in
geographic space & through geological time
Structural and functional evidence indicates
common ancestry of all ________________.
eukaryotes
Evidence that all eukaryotes share common ancestry
i. Membrane-bound organelles
ii. Linear chromosomes
iii. Genes that contain introns
What are homologous structures?
Similar structures from common ancestry
What are analogous structures?
Similar function, different ancestry (convergent evolution)
Ancestral/Derived Traits:
characteristics derived
from ancestor or from descendants
Analogous structures are due to __________ evolution. similar selective pressures result in similar phenotypic adaptations in different populations or species.
convergent
What is a phylogenetic tree?
show the amount of change over time calibrated by fossils or a molecular clock
What is a cladogram?
Shows relationships but NOT time or amount of change
What is an outgroup?
Least related lineage
nodes
represent the most recent common ancestor of any two groups or lineages.
the biological species concept provides a commonly used definition of a species for sexually reproducing organisms. it states that…
defined as a group capable of interbreeding and exchanging genetic information to produce viable, fertile offspring.
What is speciation?
Formation of new species, two populations become reproductively isolated from each other
What is allopatric speciation?
Occurs when biological populations of the same
species become isolated due to geographical
changes
What is sympatric speciation?
New species form in same location
Examples of sympatric mechanisms?
Behavioral isolation
Habitat isolation
Polyploidy
Sexual Selection
prezygotic barrier
occurs before fertilization
behavioral reproductive isolation
different mating rituals
habitat / ecological reproductive isolation
mate in different ecological environments
mechanical reproductive isolation
incompatible anatomically
temporal reproductive isolation
mate at different times of day
geographic reproductive isolation
two organisms are separated by geographical barrier
gametic reproductive isolation
gametes are unable to fuse
name the 3 postzygotic reproductive barriers
after fertilization :
Reduced hybrid viability
Reduced hybrid fertility
Hybrid breakdown
reduced hybrid viability
hybrid is not healthy / viable
reduced hybrid fertility
hybrid not fertile
hybrid breakdown
subsequent hybrid starts decreasing viability and fertility (F2)
the ______ the population, the more chance for ____________ drift.
smaller, genetic