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proximate causation
- mechanisms that produce the behavior within the animal
- what is happening in the brain that is making a fish school vs not school?
ultimate causation
- adaptive value of a behavior
- why is it beneficial for fish in one environment to school, while it is not beneficial in another environment
type of proximate causation
physiology and ontogeny
physiology
how behavior is influenced by an animal's current internal states and processes
ontogeny
how the behavior develops in an animal
types of ultimate causation
phylogeny and adaptive significance
phylogeny
why the behavior originally developed in the group of animals
adaptive significance
why is one behavior adaptive for one species but not another?
ethology
- study of animal behavior
- most ethologists use proximate causation
behavioral ecology
- study of how animals use behavior to survive and reproduce
- most behavioral ecologists use ultimate causation
associative learning
when an animal modifies its behavior because it learns to associate two stimuli or a behavior and a reward
classical conditioning
association of two stimuli (Pavlov's dogs)
operant conditioning
association of behavior and reward (Skinner boxes)
communication
when actors use signals or displays to modify the behavior of the reactors
types of communication
- acoustic
- visual
- chemical
- electric
- vibrational
- tactile (touching)
uses of communication
- mate attraction
- communication within a species
- communication across species
reproductive isolation
- prevent individuals of closely related species from mating with each other
- ex/ different species of fireflies use different light patterns and the females will only respond to the patterns of males from her own species
communication within a species
this is done to alert to the presence of food or predators, to indicate status within society, or to signal territory border
communication across species
this is done between predator and prey (prey shows predator they've seen them) or with shared vigilance between prey (prey animals warn the rest of the prey that a predator is near)
how does natural selection shape communication?
shapes displays to be as beneficial as possible; can lead to repetition, rituals, honesty, and deception
example of deceptive signal
when a blue jay imitates a hawk to drive competitors away from a food source
female mate choice
females want a few high quality mates
male mate choice
males want as many mates as possible regardless of quality
intrasexual selection
competition within one sex for the other
intersexual selection
mate choice based on desired characteristics from the opposite sex
(leads to exaggerated traits such as a peacock's tail)
what animals need to fight to win a mate
only evenly matched competitors need to fight to win a mate
sexual dimorphism
secondary sexual characteristics are traits that develop in only one of the sexes once they reach maturity that enable the to attract a mate (ie. differences between sexes)
monogamy
one male mates with one female
polygyny
one male mates with several females
polyandry
one female mates with several males
promiscuity
both males and females mate several times with different individuals
altruism
acting to increase another individual's lifetime number or offspring at a cost to one's own survival and reproduction
manipulation
recipient manipulates the donor into providing benefit (cuckoo lays egg in nest of another bird)
reciprocity
requires repeated interactions and recognition in a mutually beneficial cycle where on repays the other for help and so on
kin selection
acts that benefit blood lines thus ensuring the survival of your genes
direct fitness
fitness gained by ones own survival and reproduction
indirect fitness
fitness gained by aiding the survival and reproduction of non-descendant kin
Hamilton's rule
- c < rb
- c is cost to donor
- r is relatedness to donor and receiver
- b benefit to receiver
what does Hamilton's rule state
states that the altruistic act will be favored by natural selection as long as the cost to the donor is less than the benefit to the receiver, depending on how closely related the recipient is
aggregations
no social structure; basically just a group of animals living together
social groups
have social organizations where the individuals have stable relationships
colonies
have both social and physical structures
key stimulus/sign stimulus
a signal from the environment that triggers a stereotyped behavior
ecology
the study of how organisms interact with each other and their environment
individual adaptation (aka phenotypic plasticity)
helpful change within an individual (ex/ study schedule)(in response to unique environment)
evolutionary adaptation
inherited trait that improves the ability of individuals to survive and reproduce (ex/ cheetah's tail)
population
group of individuals of a species that are separate from other groups of that species because of factors like distance
metapopulation
group of populations that have some limited exchange between them
uniform population distribution
individuals are evenly spaced

random population distribution
result of individuals having no strong interaction with each other

clumped population distribution
usually occurs when resources are clumped

population density
the number of individuals within an area
demographics
- sex ratio
- age structure
- fecundity (birth rate per year)
- mortality (death rate per year)
generation time
average interval between the birth of an individual and the birth of its offspring
population growth rate equation
(birth rate + immigration rate) - (death rate - emigration rate)
exponential growth model

exponential growth model assumptions
assumes that there isn't any immigration or emigration and that there is no population limit
logistic growth model
has carrying capacity, or the number of individuals that can be sustained by the environment

density independent factors
- exert the same force on population regardless of the population density
- ex/ weather, catastrophes
- has negative feedback
density dependent factors
- exert greater force on growth when population is more dense
- ex/ food, predators
- has positive feedback
Allee effect
where increased population densities have increased population growth (only occurs in very small populations where its hard to find a mate
K-selected species
individuals are good survivors and competitors (humans)
R-selected species
individuals are good reproducers
rectangular population graph
even birth rate and death rate

triangular population graph
higher birth rate than death rate

inverted triangular population graph
higher death rate than birth rate
community
all the organisms that live in one place
what are communities characterized by?
they are characterized by the species within them or the species richness, species diversity, and primary productivity
ecological niche
total of all the ways that an organism uses the resources of its environment (food consumption, temperature range, moisture requirements)
fundamental niche
entire niche a species is capable of using if no other species were present
realized niche
the niche actually used by species due to the presence of other animals such as predators (deer only feed in forest because if out in the open field, predators could spot them)
two ways interspecies competition can happen
- interference competition
- exploitative competition
interference competition
two species fighting for resources
exploitative competition
one species using up all of the resources which means the second species doesn't have access to it
competitive exclusion
the inevitable elimination from a habitat of one of two different species with identical needs for resources
resource partitioning
two species can subdivide the niche by one of both of them evolving so they can coexist
resource partitioning example
when different species of lizards must share a tree, each species occupies a different area of the tree and feeds on a specific insect to allow options for the other lizards
character displacement
an advanced resource partitioning that occurs when two species evolve so they are less similar to each other; decreases niche overlap
character displacement example
Darwin's finches beaks were more similar in shape when they lived in different areas (allopatric), but were very different in shape when they lived in the same area (sympatric)
plant adaptations against herbivores
- morphological defenses
- chemical defenses
morphological defenses
thorns, spikes, small hairs, sticky secretions, toughness
evolutionary responses of herbivores to plants defenses
caterpillar evolves to be the same color as the leaf of the plant it feeds on so it is less likely to be preyed upon
animal adaptations against predators
- secondary compounds in diet
- chemical defenses
animal use of secondary compounds in diet
ex/ monarch butterfly eats milkweed so its body is filled with toxins to avert predators from eating it
animal use of chemical defenses
ex/ poison dart frogs
- warning coloration: bright colors and patterns
batesian mimicry
organism without defensive chemicals resembles a species that has them

mullerian mimicry
when two or more species with defensive chemicals look similar to one another

symbiosis
species that have evolved to live together
mutualism
both species benefit from interaction
parasitism
one species benefits, the other is harmed
three kinds of parasites
- endoparasites (internal - worms)
- ectoparasites (external - ticks)
- parasitoids: lay their eggs in or on other organisms so their young can hatch and feed on the host
commensalism
one species benefits, the other is not affected
effects of predation and parasitism
reduced competition in prey species by lowering the number of individuals competing for a resource
what is an indirect effect?
when two species affect each other through another species
ex/ presence of rodents decreases the amount of large seeds, which increases the amount of small seed for ants to feed on
keystone species
species whose effect on the the composition of communities are greater than expected for the abundance of said species (beavers)
ecological succession
tendency of communities to change from simple to complex over time
primary succession
when a community develops from nothing (ex/ where volcanoes lay down new rock)
secondary succession
when a community is disturbed but organisms remain
three components of an ecosystem
- non living things
- producers
- consumers
the water cycle steps
1. evaporation
2. condensation
3. precipitation
4. transpiration
5. groundwater
