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Osmoregulation
is the maintenance of constant [solute] in the body fluids
Osmoregulatory
______ demands vary across environments
excretion
Osmoregulation is linked to _____ of metabolic wastes
Vertebrate kidney
produces waste fluid and regulates blood [solute]
Solute concentration
_______ of the body fluids is critical to cell function
solutes
are substances dissolved in fluid
ex: salt
Gradients in [solute]
what dictate water movement across cell membranes
[Solute]
_____ means solute concentration
osmosis
______ is movement of water from an area of low [solute] to an area of high [solute] Water Balance & Solute Concentration
isoosmotic environment:
fluid [solute] = cell [solute]
Water moves into & out of the cell at equal rates
• Cell does not gain or lose water
hypoosmotic environment:
fluid [solute] < cell [solute]
Water moves into the cell
• Cell can burst under pressure
hyperosmotic environment:
fluid [solute] > cell [solute]
Water moves out of the cell
• Cell can shrivel and die
Marine animals
high [solute] environment example organism
Freshwater animals
low [solute] environment example organism
Terrestrial animals
dehydrating environment example organism
osmoconformers:
• Body fluid [solute] = sea water [solute]
Many marine inverts are
osmoregulators:
• Body fluid [solute] < sea water [solute]
• Sea water is dehydrating!
• Tend to lose water and body fluid becomes too
concentrated
Produce a concentrated waste fluid to retain water
and shed [solutes]
Many marine vertebrates are
osmoregulators:
Body fluid [solute] > freshwater [solute]
• Body fluid tends to become too dilute
• Produce a dilute waste fluid to expel water and
retain [solutes]
Freshwater animals are
osmoregulators
• Water loss → increase body fluid [solute]
• Produce a concentrated waste fluid to retain water
and shed solutes
(Terrestrial animals lose water by evaporation)
Terrestrial animals are
Amount & type of waste affects water balance
Example
• Nitrogenous waste is particularly important
• Metabolism of amino acids & nucleic acids produces
ammonia (NH3)
• NH3 is toxic
• Forms NH4+ → interferes with ATP synthesis
• Ammonia can be excreted directly but must be diluted
• Can be converted to less toxic forms, but this costs energy
Animals that face water limitation convert ammonia to____
urea or uric acid
• Urea is more energetically expensive than ammonia
but requires less dilution
• Uric acid is more energetically expensive than urea &
ammonia and requires almost no dilution
Excretory systems
remove nitrogenous wastes
and perform osmoregulatory functions
• Collect nitrogenous waste from the body fluid
• Control the amount of water excreted with the
waste fluid
• Pump solutes in or out of waste fluid depending on
body fluid [solute]
Excretory System Function
Produce a waste fluid by refining a filtrate derived from blood/body fluid
1. Filtration - nonselective movement of body fluid into excretory tubule
2. Reabsorption – pump useful molecules back into blood
3. Secretion – pump toxins, waste, and unwanted solutes into filtrate
4. Excretion – release waste fluid (urine)
Excretory systems are
networks of tubules
Tubules separate
stages of waste processing
SA
Packing of tubules increases
higher rate of waste fluid processing
Increased SA allows for
Transport epithelium
Tubules are lined by
Transport epithelium
is made up of specialized cells that move particular solutes in specific directions
Kidneys
paired organs each made up of networks of highly folded tubules
Renal artery
brings blood to kidney
Renal vein
drains blood from kidney
ureter
Urine produced in the kidneys exits through the
urinary bladder
Ureter drains to the
urethra
Urine is expelled through the
renal cortex
Kidney has an outer
renal medulla
Kidney has an inner
Renal pelvis
collects waste fluid and drains to ureter
nephrons
Cortex and medulla are packed with ____ (excretory tubules)
blood vessels
Nephrons are associated with
Nephrons
collect blood filtrate and exchange materials with blood in capillaries
Bowman’s capsule
Filtration happens in what part of the Nephrons
glomerulus
Blood pressure causes fluid to leak out of the
glomerulus
is capillary bed housed within Bowman’s capsule
nonselective filtering
what type of filtering occurs in the Bowman’s capsule
Proximal tubule
Loop of Henle
Distal tubule
What three regions of nephron does processing of waste fluid happen in?
Descending arm of Loop of Henle
reabsorbs water from waste fluid
Ascending arm of Loop of Henle
pumps salt out of waste fluid
Waste fluid drains from distal tubule into
collecting duct
Proximal tubule
Reabsorbs water, nutrients, & some salts
Distal tubule
allows for more exchange of salts
collecting ducts
Water can be reabsorbed from waste fluid in
hormonal control
Collecting duct’s permeability to water is under