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what phylum has the least constraint of body plan
phylum chordata
two invertebrate subphyla of phylum chordata
cephalochordata and urochordata
what developmental patterns does phylum chordata have
deuterstome
what two phyla are chordates related to
echinoderms and hemichordates
what date did phylum chordata originate
precambrian origin with rapid diversification in the paleozoic
5 distinctive features of phylum chordata
notochord, hollow dorsal nerve cord, pharyngeal gill slits, endostyle, postanal tail
notochord
endoskeleton for muscles; lost or replaced in some
hollow dorsal nerve cord
controls muscles, includes brain; lost or replaced in some
pharyngeal gill slits
open to outside; respiration and feeding; become other structures in vertebrates
endostyle
generates mucus for feeding; thyroid gland in vertebrates
subphylum cephalochordata
our primitive cousins; consists of lancelets or “amphioxus”; “little fish without heads”; all 5 chordate features present
how do cephalochordata feed
filter feeds by passing water over pharyngeal slits covered in mucus; enters though gill slits to the atrium and out the atriopore
circulatory system for subphylum cephalochordata
closed blood vascular system; no heart (peristalsis of vessels); no blood pigments or cells
reproductive system for subphylum cephalochordata
simple gonads; gametes shed directly into atrium and out atriopore
subphylum urochordata
our veyr weird cousins; loss of all chordate structures
two classes we need to know for subphylum urochordata
class ascidiacea (sea squirts) and class appenduclaria (planktonic)
class ascidiacea
sea squirts; solitary and colonial as adults; tadpole larva has 5 chordate features; filter feed; when colonial, they are zooids
class appendicularia
abandoned houses major source of marine snow; important for global carbon cycling
post-anal tail
propulsion, often with fins; vestigial in humans (coccyx)
subphylum vertebrata of phylum chordata
has vertebral column, cranium, endoskeleton, neural crest
vertebral column
notochord replaced by column of interlocking vertebrae
cranium
protective housing around the brain
endoskeleton
grows with body; cartilage, then-bone
neural crest
embryonic cells, contribute to development of cranium
5 extant classes of subphylum vertebrata
myxini, petromyzontida, chondrichthyes, actinopterygii, sarcopterygii
myxini
hagfishes; marine scavengers; have slime glands to make sheets of mucus
petromyzontida
lampreys; marine and freshwater
chondrichthyes
sharks, skates, chimaeras; mostly predators; most are marine; no swim bladder
actinopterygii
ray-finned fishes
sarcopterygii
lobe-finned fishes
how did the evolution of the endoskeleton progress
notochord to vertebral column, cartilage to bone
how did the evolution of the pharynx progress
pharyngeal slits to gills, greater respiratory capacity
how did the evolution of the nervous system progress
greater development of senses and brain
how did the evolution of the appendages progress
greater development of paired pectoral and pelvic fins
how did the evolution of the jaws progress
modified cartilaginous gill arches derives from neural crest cells with jaw muscles from gills
what do myxini lack
lack jaws, scales, paired fins, bone, and vertebral column
what are teeth of myxini made of and used for
made of keratin on tongue; used for rasping food
what do petromyzontida lack
lack jaws, scales, paired fins, and bone
what are teeth of petromyzontida made of and used for
made of keratin around sucker-like mouth; may be used to parasitize
what does the life cycle involve for class petromyzontida
involves ammocoetes larva that filter-feeds and looks like a cephalochordate (lancelet)
what do chondrichthyes have
have jaws, paired pelvic and pectoral fins, placoid scales adapted to form teeth
what kind of skeleton do chondrichthyes have
cartilaginous skeleton
how do chondrichthyes maintain balance between water and salt
osmoregulation; requires copulation and shelled eggs
what does class actinopterygii have
have jaws, paired pelvic and pectoral fins, light-weight cycloid, ctenoid or ganoid scales, swim bladder
what kind of skeleton does actinopterygii have
bony skeleton (endochondral); gills are covered by bony operculum
what does osmoregulation support in actinopterygii
permits free-spawning and planktonic development
what does class sarcopterygii have
have jaws, paired pelvic and pectoral fins, heavy enameled scales
what kind of skeleton does sarcopterygii have
bony skeleton
what does the swim bladder do in sarcopterygii
acts as a lung in lungfishes
what did sarcopterygii give rise to
tetrapods
what do sarcopterygii look like
they look like the “living fossil” Coelocanth, a mesozoic relic