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Dessication
Excessive water loss; drying out
Desiccation tolerance
A suite of biochemical traits that allow cells to survive extreme dehydration by minimizing damage to membranes and macromolecules
Shoot
The collective name for the leaves, stems, and reproductive organs; the major above ground organ systems of vascular plants
Epidermis
The outermost layer of cells in multicellular organisms such as plants and animals
Vascular Tissue
A plant tissue that extends from leaves to roots and allows water and nutrients to move by bulk flow
Ground Tissue
Everything that is neither epidermis nor vascular tissues
Parenchyma
Thin-walled cells that perform many metabolic functions, including photosynthesis and storage of nutrients, carbohydrates, and water
Mesophyll
The photosynthetic cells that make up the tissue between the upper and lower epidermis of plant leaves
Vein
A vascular conduit within the leaf; in animals, a large, low pressure vessel that returns blood to the heart
Transpiration
The loss of water from leaves
Guard Cell
One of two cells surrounding the central pore of a stoma and that regulates the opening and closing of this pore
Crassulacean Acid Metabolism (CAM)
A process in plants that helps balance carbon dioxide gain and water loss by opening stomata and capturing carbon dioxide into 4-carbon organic acids at night, when transpiration rates are low, and then using it to supply the Calvin cycle during the day while stomata remain closed
C4 Plant
A plant in which carbon dioxide is incorporated into 4-carbon organic acids in mesophyll cells that are then used to supply the Calvin cycle in bundle-sheath cells; results in the suppression of photorespiration
C3 Plant
A plant in which carbon dioxide is first incorporated into a 3-carbon molecule during photosynthesis. This contrasts with a plant, in which carbon dioxide is first incorporated into a 4-carbon molecule
Vessel Element
An individual cell that is part of a xylem vessel for water transport in plants
Pit
In vascular plants, porous regions that allow water to flow from one xylem conduit to another, but prevent air from spreading
Water Potential
The free energy of water in which water moves from an area of high water potential to an area of low water potential
Cavitation
The abrupt replacement of liquid water in a xylem conduit by water vapor, which prevents water from being pulled through the xylem by transpiration
Source
Regions that produce or store carbohydrates
Sink
Any portion of the plant that needs carbohydrates to fuel growth and respiration; examples are roots, young leaves, and developing fruits
Companion Cell
In angiosperms, a cell associated with sieve tubes that carries out cellular functions such as protein synthesis.
Phloem Sap
The sugar-rich solution in plants that flows through both the lumen of the sieve tubes and the sieve plate pores
Sieve Tube
In angiosperm phloem, a multicellular unit composed of sieve elements that are connected end to end, through which phloem transport takes place
Sieve Plate
A modified end wall with large pores that links sieve elements
Rhizosphere
The soil layer that surrounds actively growing roots
Root Hairs
A slender outgrowth produced by epidermal cells that greatly increases the surface area of the root.
Endodermis
A layer of cells that acts as a gatekeeper controlling which nutrients move into the xylem
Casparian Strip
A thin band of hydrophobic material that prevents water and any dissolved materials from moving through the cell wall
Mycorrhizae
Symbioses between roots and fungi: the plant provides carbohydrates produced by photosynthesis, and the fungus provides nutrients it has obtained from the soil
Ectomycorrhizae
One of the two main types of mycorrhizae; ectomycorrhizae produce a thick sheath of fungal cells (hyphae) that surround the root tip, as well as hyphae that grow between, but do not penetrate, cells in the interior of the root
Endomycorrhizae
One of the two main types of mycorrhizae; endomycorrhizal hyphae penetrate into root cells, where they produce highly branched structures (arbuscules) that provide a large surface area for nutrient exchange
Nitrogen Fixation
The process by which some Bacteria and Archaea convert nitrogen gas into ammonia, a form biologically useful to primary producers
Root Nodule
A structure, formed by dividing root cells, in which nitrogen-fixing bacteria live
Shoot apical meristem
A group of totipotent cells at the tip of a stem or branch that gives rise to new shoot tissues in plants.
Axillary bud
A meristem that forms at the junction between a leaf and the stem
Node
A point along the stem where one or more leaves are attached and which has one or more axillary buds
Internode
The section of stem between successive nodes
Meristem identity gene
A gene that contributes to meristem stability and function
Petiole
A structure that connects a leaf to its stem
Bud scale
Small modified leaves that protect shoot apical meristems from desiccation and damage due to cold
Primordia
An organ in its earliest stage of development; in plants, leaf primordia form near the tips of shoot apical meristems and develop into leaves
Procambial cell
A plant cell that retains the capacity for cell division and gives rise to both xylem and phloem
Cortex
The tissue between the epidermis and the vascular bundles
Pith
In a stem, the region inside the ring of vascular bundles
Root apical meristem
A group of totipotent cells near the tip of a root that is the source of new root cells
Root cap
A structure that covers and protects the root apical meristem as it grows through the soil
Pericycle
In roots, a single layer of cells just to the inside of the endodermis from which new root apical meristems are formed, allowing roots to branch
Lateral meristem
The source of new cells that allows plants to grow in diameter
Vascular cambium
Lateral meristem that is the source of new xylem and phloem
Cork cambium
Lateral meristem that renews and maintains an outer layer that protects the stem against herbivores, mechanical damage, desiccation, and fire
Secondary xylem
New xylem cells produced by vascular cambium, which are located to the inside of the vascular cambium
Secondary phloem
New phloem cells produced by the vascular cambium, which are located to the outside of the vascular cambium
Sapwood
In long-lived trees, the layer adjacent to the vascular cambium that contains the functional xylem
Heartwood
The center of the stem in long-lived trees, which does not conduct water
Growth ring
One of the many rings apparent in the cross section of the trunk of a tree, produced by decreases in the size of secondary xylem cells at the end of the growing season, that make it possible to determine the tree's age
Suberin
In plants, a waxy compound found in the walls of cork cells that protects against mechanical damage, the entry of pathogens, and water loss
Lenticel
Small regions where the outer bark cells are less tightly packed
Auxin
A plant hormone that plays a major role in plant development through the establishment of concentration gradients that guide patterns of cell growth and differentiation
Polar transport
The coordinated movement of auxin across many cells
Gibberellin
A plant hormone that stimulates the elongation of stems
Cytokinin
A plant hormone that stimulates cell division and delays leaf senescence
Apical dominance
The suppression of growth of axillary buds by the shoot apical meristem, typically by the action of auxin hormone
Strigolactone
A plant hormone that inhibits the outgrowth of axillary buds
Tropism
The bending or turning of an organism in response to an external signal such as light or gravity
Phototropic
When plants bend toward the light
Gravitropic
When plants grow upward against the force of gravity
Statolith
Specialized gravity-sensing cells in the root cap contain large starch-filled organelles
Phytochrome
A photoreceptor that switches back and forth between two stable forms, active and inactive, depending on its exposure to red and far-red light
Abscisic acid
A plant hormone that triggers stomatal closure, stimulates root elongation, and maintains seed dormancy
Thigmotropism
Developmental response to a mechanical stimulus arising from physical contact
Photoperiodism
The effect of the photoperiod, or day length, on physiological or developmental processes; in plants, the effect of day length on flowering
Short-day plan
A plant that flowers only when the day length is less than a critical value
Long-day plant
A plant that flowers only when the duration of the light period exceeds a critical value
Day-neutral plant
A plant that flowers independently of any change in day length
Vernalization
A prolonged period of exposure to cold temperatures necessary to induce flowering in some plants
Host plant
Species that can be infected by a particular pathogen
Biotrophic pathogen
A plant pathogen that obtains resources from living cells
Necrotrophic pathogen
A plant pathogen that kills cells before drawing resources from them
Virulent
Describes a microbe or virus that causes disease or significant damage to the host
Avirulent
Describes a microbe or virus that lacks the ability to cause disease or significant damage to the host
Basal resistance
A component of the plant immune system that recognizes molecules produced by broad classes of pathogens; contrast with specific resistance
Specific resistance
A component of the plant immune system that recognizes proteins produced by specific pathogens; contrast with basal resistance
Avr protein
An avirulence protein produced by a plant pathogen that blocks the host plant's immune response and is detected by resistance (R) proteins
R protein
Any one of a group of resistance proteins in plant cells, each expressed by a different R gene, that functions as part of the plant's immune system by detecting the presence of a specific pathogen-derived Avr protein
R gene
Any one of a group of genes that expresses an R (resistance) protein in plants
Hypersensitive response
A type of plant defense against infection in which uninfected cells surrounding the site of infection rapidly produce large numbers of reactive oxygen species, triggering cell wall reinforcement and causing the cells to die, thus creating a barrier of dead tissue
Systemic acquired resistance (SAR)
The ability of a plant to resist future infections, occurring in response to a wide range of pathogens
Small interfering RNA (siRNA)
A type of small double-stranded regulatory RNA that becomes part of a complex able to cleave and destroy single-stranded RNA with a complementary sequence
Ti plasmid
A small circular DNA molecule in virulent strains of Agrobacterium tumefaciens containing genes that can be integrated into the host cell's genome, as well as the genes needed to make this transfer
Trichome
Hairlike outgrowths on the surfaces of leaves and stems; may provide protection from herbivores and excess sunlight
Latex
A white sticky liquid produced in some plants
Alkaloid
Nitrogen bearing compounds that damage the nervous system of animals
Terpene
Any one of a group of compounds that do not contain nitrogen and are produced by some plants as a defensive mechanism
Tannin
Any one of a group of phenols found widely in plant tissues that bind with proteins and reduce their digestibility
Protease inhibitor
A molecule that blocks the action of enzymes that break down proteins
Extrafloral nectar
A nectar-producing gland not associated with flowers, often located on leaves
Constitutive Defense
Defenses that are always present
Inducible defense
A defense that is activated only when the plant senses the threat
Integrated pest Management (IPM)
A method of pest control that monitors pest populations, encourages predator populations, applies mechanical and targeted chemical control when pest populations reach levels that cause economic losses, and uses nonspecific pesticides only as a last resort
Porifera
Sponges; an animal phylum characterized by few cell types, no well-defined tissues, and no clearly defined plane of symmetry