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Ecdysozoans are _______ along with lophotrochozoans
Protostomes
The defining characteristic of ecdysozoans is
Ecdysis
Ecdysis is?
molting a cuticle
Phylum Nematoda
Microscopic round worms, hydrostatic skeleton, largest number of individuals on the planet
Nematodes are constrained because the blastocoel…?
Is filled with hydrostatic pressure and becomes the pseudocoel
Nematode parasites are different from lophotrochozoan parasites because they lack?
Tegument, they must feed via the mouth
Guinea worms
A species of nematode, parasitic, contracted from eating contaminated eggs or water, female worm emerges from skin to spread eggs
Phylum Arthropoda is considered the ____ phylum because it has ______
Largest phylum (1 million species)
Exoskeleton causes arthropods to be ?
Moderately constrained
Exoskeleton
Thick and hard cuticle made of protein, limestone, and chitin
Growth in arthropods is
Discrete- ecdysis
Apodernes
Allows flexion of the jointed limbs, contracts against exoskeleton
Metameric body
Body regions that are specialized
Arthropods have a _____ blood vascular system
Open
Four subphylum of arthropods
Chelicerata, Myriapoda, Crustacea, and Hexapoda (Insecta)
Subphylum Chelicerata
Defined by lack of mandibles, antennae, and external sex organs
What do chelicerates use in place of mandibles?
Chelicerae fangs
Chelicerata have _____ legs and one extra pair called ______
Four walking legs, pedipalps
The three classes of chelicerata are ?
Class merostomata, pycnogonida, and arachnida
Class Merostomata
Horseshoe crabs, living fossil, telson to flip self over with
Class Pycnogonida
Sea spiders, digestive glands + gonads in legs
Class Arachnida
Terrestrial, extracorporeal digestion
The most successful arachnid is ____ because they produce _____
Spiders, webs
Class myriapoda
Head and many segments, no tagmata
Two classes of Myriapods
Class diplopoda and chilopoda
Class diplopoda is known as
Millipedes
Class chilopoda is known as
Centipedes
Diplopod key traits:
Herbivores, slow, thick cuticle, 2 legs per segment
Chilopod key traits:
Predators, fast, thin cuticle, one leg per segment, fangs
Subphylum Crustacea
Biramous appendages, mandibles, two tagmata- Cephalothorax and abdomen, Nauplius larva
Class insecta
Greatest number of described species, uniramous appendages, malpighian tubules, huge land abundance
Body plan of class insecta
Head, thorax, and abdomen
The head of an insect must include ?
Antennae, compound eyes, and mandibles
Respiration in class insecta
Breath via tracheids that flow directly into the muscles, bidirectional spiricule takes in oxygen by diffusion
Malpighian tubules
Excretory system in insects, excrete uric acid and dry waste to conserve water
Flight in insects
Possible due to bistability in exoskeleton, 86% efficiency, non-living structural exoskeleton controlled by longitudinal/indirect flight muscles
Indirect flight muscles contract to make the thorax _____ which makes the wings _____
Flip inwards; go up
Longitudinal muscles contract to make the thorax _____ which makes the wings
Flip back outwards; go down
Sociobiology
The relationship between genetics and behavior
Eusocial behavior in insects like bees or ants is caused by?
Haploid sex determination
How closely related are male bees to the queen bee?
they are direct clones
How closely related are sister bees to each other?
They are more genetically alike (3/4) than if they were to have their own offspring (1/2)
Organisms such as bees ultimately engage in social behavior because they want to spread their …?
Genes
Holometabolous life history
Derived insect life, egg > maggot > pupa > adult
Superphylum Deuterostomia includes all except?
Ecdysozoans
What is the defining characteristic of Echinodermata?
Pentaradial symmetry
Aboral surface means the ?
Top, dorsal side, contains anus
Oral surface means?
Bottom, ventral side, contains mouth
Water vascular system
water moves by internal ampullae, pushes external tube feet for locomotion, also used in respiration
Nervous system in echinoderms
Controls ampullae and tube feet, found directly under vascular system
Pedicellaria
Provides protection of the echinoderm body and can release toxins in some species
Perivisceral coelom
Space in echinoderms, holds the two major organ groups
Pyloric cecum
Cardiac stomach for extracorporeal digestion (echinoderms)
How do grazing echinoderms feed?
Aristotle’s lantern
Echinoderm reproduction is interesting because the juvenile ____ the larva that produced it
Eats
What phylum has the least constrained body plan?
Chordata
Three subphyla of Chordata?
Cephalochordata, Urochordata, and Vertebrata
Notochord
endoskeleton for muscles, protects the dorsal nerve cord, becomes boney in vertebrates
Dorsal Hollow Nerve Cord
Controls muscles, includes brain, becomes spinal cord in vertebrates
Pharyngeal gill slits
Open to outside, feeding and respiration, become other structures in vertebrates
Endostyle
Generates mucus for feeding, thyroid gland in vertebrates
Post-anal tail
Used for propulsion, vestigial structure in humans (tailbone)
Lancelets
Subphylum Cephalochordata
What is the defining characteristic of subphylum cephalochordata?
Adults retain all 5 features of a chordate
How do cephalochordates feed?
Filter feeding by passing water over pharyngeal gill slits
Reproduction in cephalochordata
Segmented gonads, Gametes shed directly into atrium and out atriopore
What is the defining characteristic of subphylum Urochordata?
Loss of adult chordate features and greater filter feeding adaptations
Three classes under urochordata
Class Ascidiacea, Class Thaliacea, and Class Appendicularia
Class Ascidiacea
Sea squirts, can be colonial, tadpole larva with all 5 chordate features, sessile as adults
What two features do urochordates retain as adults?
Pharyngeal gill slits and endostyle for feeding
How is class ascidiacea useful for man?
Produce antitumor agent Yondelis
Class Thaliacea
Planktonic water columns, can be colonial, 3 orders with complex life histories, can grow very large and quick
Class Appendicularia
Planktonic, retain all chordate features, build tiny mucus houses that contribute to the carbon cycle
Protandric hermaphrodites
Class Appendicularia, die when giving birth
Marine snow
Mucus houses that fall to the bottom of the ocean after Appendicularia has used it
Subphylum Vertebrata
Vertebral column, Cranium, Neural crest, and endoskeleton
Vertebral column
notochord replaced by column of interlocking vertebrae
Neural crest
embryonic cells, contribute to development of the cranium
How many extant classes are there in vertebrata?
5
How did the vertebrate evolve?
Notochord to vertebrae, bones, greater respiratory and nervous system, paired pectoral and posterior fins, development of jaws
How did Jaws develop?
Modified cartilaginous gill arches (numbers 3 and 4)
Class Myxini
Hagfish, Lack jaws, scavengers (eat dead whales), secret mucus as defense
Class petromyzontida
Lampreys, Lack jaws, sucker mouth and many teeth, ammocoetes larva
Class Chondrichthyes
Sharks and rays, Has jaws, Cartilaginous, pectoral fins, placoid scales adapted to form teeth, no swim bladder
Chondrichthyes reproduction
Osmoregulation requires copulation and shelled eggs
Class Actinopterygii
Ray finned fish, Has jaws, Homocercal caudal fin, operculum coverers gills, swim bladder
What makes Class Actinopterygii unique?
Posterior fin without bones, adapted to all aquatic environments
Boney fish include?
Class actinopterygii and sarcopterygii, as well as the tetrapods
Boney fish reproduction differs from cartilaginous fish because?
Osmoregulation permits free-spawning and planktonic development
Class Sarcopterygii
Lobe-finned fishes, 8 species, has jaws, Diphycercal caudal fin - Strong, fleshy lobed fins – lineage gave rise to tetrapods
Lungfish
Evolved swim bladder, living fossil- Coelacanth species
What evolution causes tetrapods?
Pectoral and pelvic girdles and limbs originated for balance in swimming but aided on movement to land
When did vertebrates make it to land and why?
Devonian period- 400 mya, vascular plants, molluscs, and arthropods available to eat
Important adaptations for land dwelling
Limbs and lungs, temperature regulation and prevention of desiccation
Why are lungs needed?
Oxygen content higher in air than water
Tetrapod classes
Class amphibia, reptilia, and mammalia
Class Amphibia
Three suborders - Salamanders, frogs, and caecilians, 4 limbs, Lungs with internal nostrils, Metamorphosis of aquatic
larvae to terrestrial adult, Ectothermic
What is unique about amphibian sensory system?
Lateral-line system
Amphibian brain development
Forebrain for olfaction, midbrain for vision, and hindbrain for hearing and balance
Vascular system in amphibians
3-chambered heart- Mixed circulation where oxygenated blood returns to heart and mixes with deoxygenated blood