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what is differentiation
the process by which cells become different from one another
how do plant cells increase in size
by taking in what in what is called the stress-relaxation cycle
how does the stress-relaxation cycle work
the cycle begins when turgor pressure acts on the cell wall. This physical stress alters the structure of the material that makes up the cell wall. Some changes are reversible, some arent and will become permanently stretched out. Then the wall relaxes, turgor pressure falls and the cycle repeats.
what can change the rate of cell wall expansion
by changing the chemical composition of the wall. Some plant hormones increase the rate of cell expansion by making the cell wall looser.
what dictates the direction of maximal cell expanion
it usually can be predicted by the arrangement of cortical microtubules (cytoskeletal proteins that are anchored to the cell membrane)
what is a dormant seed
when the development of a seed stops even when external conditions are adequate for development
how do seeds maintain dormancy
exclusion of water or oxygen from the embryo
mechanical restraint of the embryo by a tough seed coat
chemical inhibition of germination by growth regulators
photodormancy: some seeds need a period of light or dark before they can germinate
thermodormancy: some seeds need high or low termperature to germinate
why do seeds become dormant
dormancy ensures that the seed will germinate at a time suitable for the plant to complete its life cycle
some seeds need the cold of winter to germinate in the spring
when do seeds germinate
seeds begin to germinate or sprout when dormancy is broken and environmental conditions are satisfactory.
how does germination start
imbibition. The seed takes in water because of the polar macromolecules, like cellulose in the seed. Seeds also take in water because of their very negative water protential
process of germination
starch, proteins, and lipids are hydrolyzed to provide energy and building blocks for the growing embryo. Reserves are stored in cotyledons. Germination is complete when radicle emerges from seed coat.
what is a cotyledon
the first leaf or leaves of the embryo
what is a radicle
embryonic root
what do photoreceptors allow to happen
when seeds germinate underground they need to know which way is up. Photoreceptors that sense light and gravity direct this stage of development
what is the coleoptile
in monocots its a sheath of cells which protects the shoot as it grows and pushes its way through the soil

what are photoreceptors
they detect changes in the quality and direction of light as well as the timing of light availability.
what are tropisms
a directional response to a stimulus
what is phototropism
the growth or movement of a plant towards light
what is gravitropism
the growth of a plant towards gravity
what is thigmotropism
the growth of a plant in response to touch
what are plant hormones
chemical signals in very low concentrations who play multiple regulatory roles at target sites often distinct from where they are produced.
effects of gibberellins on plant growth and development
stem elongation
fruit growth
mobilization of seed reserves
where is auxin produced
in the shoot apex and diffused down the shoot in a polar fashion
main roles of auxin
plant growth and development
what does the polar transport of auxin depend on
diffusion across a cell membrane
membrnae protein asymmetry
proton pumping/chemiosmosis
ionization of a weak acid
explain how the polar transport of auxin depends on the diffusion across a cell membrane
polar molecules diffuse across cell membranes less readily than nonpolar molecules
explain how the polar transport of auxin depends on membrane protein asymmetry
some proteins that transport auxin are located in specific portions of the cell membrane. In the stem, most of the transport proteins that allow auxin to move out of the cell are located at the bottom end of the cell
explain how the polar transport of auxin depends on proton pumping/ chemiosmosis
a proton pump moves protons from the cytoplasm to the cell wall, thereby increasing the intracellular pH and decreasing the pH in the cell wall.
Proton pumping also sets up an electrochemical gradient, which provides potential energy to drive auxin transport
explain how the polar transport of auxin depends on ionization of a weak acid
the main form of auxin, indole-3-acetic acid, is a weak acid. When the pH is low, this reaction is driven to the right, and HA (non-ionized auxin) is the predominant form. When the pH is higher, there is more A- (ionized auxin)

how are polar auxin transporters distributed
they are distributed such that the hormone predominately moves down the axis of the shoot. However, lateral redistribution of auxin is responsible for directional plant growth
what is step 1 of polar auxin transport
auxin enters the cell by passive diffusion as an unchanged acid or through auxin import proteins, symported with proton.

what is step 2 of polar auxin transport
proton pumps in the cell membrane maintain the cell wall at an acidic pH and set up a chemiosmotic gradient to drive the transport of auxin

what is step 3 of polar auxin transport
auxin- predominates in the cytosol, which has a neutral pH

what is step 4 of polar auxin transport
auxin- exits the cell via auxin anion efflux(move out of cell) carriers that are concentrated at the bottom end of each cell

what is step 5 of polar auxin transport
in the cell wall, the lower pH causes auxin- to become auxin, which can diffuse into the next cell or enter through an auxin-proton symporter

why do plants bend toward the light
the lateral movement of auxins toward the shaded side within the tip, this will stimulate cell elongation on the shaded side as the auxin travels down, causing the coleoptile to grow toward the light

auxin’s role in leaf abscission
leafs normally detach in a process called abscission, but auxin prevents this.
auxin’s role in root initiation
serves as the primary chemical signal for root initiation. They dictate where and when new roots form
auxin’s role in apical dominance
auxin helps maintain apical dominance
what is apical dominance
its where apical (terminal) buds inhibit the growth of axillary buds, resulting in the growth of a single main stem with minimal branching
auxin’s role in fruit development
fruit development normally needs prior fertilization, but auxins causes formation without fertiliation
auxin’s role in cell expansion
auxin regulates the cell wall expansion
what is cell expansion driven by
the uptake of water, which enters the cytoplasm of the cell and accumulates in its central vacuole.
explain the acid growth hypothesis
auxin increases the pumping of protons from the cytoplasm into the cell wall, this lowers the pH of the wall and activates the enzyme expansins which loosen the cell wall, making it easier to stretch as the cell expands.

what are the two aspects of light
its quality, the wavelengths of light that can be absorbed by molecules in the plant
its quantity, the intensity and duration of light exposure
which color is most effective in inducing the coleoptiles to curve
blue light
What is phototropin
a blue-light receptor protein which absorbs blue light
what is cryptochromes
it is a class of blue-light receptors which absorb blue and ultraviolet light
list some of the red light induced changes
the apical hook straightens, the rudimentary leaves unfold, and chlorophyll is made
key feature of red light responses
they are reversible by far-red light (710 - 740nm)
what are phytochromes
red light receptors
how do phytochromes switch between states
the default state, which absorbs red light, is Pr. When Pr absorbs a photon of red light it is converted into Pfr
What is Pfr
the active form of phytochrome - the form that triggers important biological processes in various plants, like chlorophyll synthesis
what does the chromophore of Pr do
the chromophore of Pr preferentially absorbs red light; when it does so, it changes conformation and the phytochrome is converted to the Pfr form.
what part of phytochrome absorbs red and far-red light
its a covalently attached pigment called a chromophore
what happens to the chromophore of Pfr when it absorbs far-red light
when the chromophore of Pfr absorbs far-red light, hte phytochrome is converted back to the Pr form
what is dark reversion
Pfr form can slowly revert to the Pr form. This even occurs gradually and doesn’t not require light
what happens to plants growing in the shade
in the shade of other plants, the ratio of red to far-red light is low (0.13:1) which stimulates cell elongation and the plants grow upward toward the sunlight
what is etiolation
To outcompete neighbors for sunlight, the plant directs energy into rapid stem elongation and upward growth rather than branching, resulting in tall, thin plants (etiolation
how does the conversion from Pr form to Pfr form actually happen
when Pr absorbs red light, the chromophore changes shape, causing change in the conformation of the protein itself, from Pr to Pfr. This results in the movement of Pfr from cytoplasm to nucleus, once there it can stimulate genes that were repressed and repress genes that were formerly stimulated.

what is photosynthesis
an anabolic process by which energy of sunlight is captured and used to convert CO2 into more complex carbon-containing compounds
what is the chemical reaction of oxygenic photosynthesis
6CO2 + 6H2O > C6H12O6 + 6 O2

what are the two pathways of photosynthesis
the light reactions
the light-independent reactions
what are light reactions
they convert light energy into chemical energy in the form of ATP and NADPH.
-NADPH acts as a reducing agent in photosynthesis and other anabolic reactions
what are light-independent reactions
they do not use light directly, but instead use the ATP and NADPH made by the light reactions and CO2 to produce carbohydrate
what can happen when a photon meets a molecule
it can be reflected
it can be transmitted
it can be absorbed, adding energy to the molecule
what happens when a molecule absorbs a photon
the molecules acquires the energy of the photon. The increase of energy boosts one of the electrons in the molecule into a shell far from its nucleus, the electron here is held less firmly, making the molecule more unstable and more chemically reactive
what are pigments
molecules that absorb wavelengths in the visible spectrum
why do leaves appear green
when white light hits the leaf, chlorophyll absorbs both blue and red light, and transmits green light which we are able to see
what is an absorption spectrum
a graph that plots light absorbed by a pigment against wavelength. its used to identify light capturing properties of molecules

what is an action spectrum
a graph that plots the rate of a biological activity carried out by an organism against the wavelength of light to which it is exposed.

what is a photosystem
an entire complex comprised of a long hydrocarbon tail which anchors chlorophyll a molecule to a complex of proteins that span the thylakoid membrane as well as other pigments like chlorophyll b, cartenoids, and phycobilins.
what are the macromolecules
carbon, hydrogen, oxygen, and nitrogen
aside from being a source of energy what does fixed carbon molecules from photosynthesis form
the physical building blocks from which the plant body is constructed.
where do plants obtain most of their nutrients
from the soil
what is a soil solution
within the soil, minerals dissolve in water as ions, forming a solution that contacts the roots of plants
what is an essential element
a plant nutrient that if the element is absent of insufficient the plant fails to complete its life cycle or grows abnormally
what are the two types of essential elements in plants
macronutrients
micronutrients
how much macronutrients does a plant need
a plant needs macronutrients in concentrations of at least 1 gram per kilogram of the plant’s dry matter
how much micronutrients does a plant need
a plant needs micronutrients in concentrations of less than 100 milligrams per kilograms of the plat’s dry matter
mineral elements required by plants (just a chart don’t memorize this)

what is a deficiency symptom
when a plant is deficient is an essential element, it displays characteristic symptoms
how are essential elements identified in plants
plants were grown hydroponically (in a nutrient solution instead of soil) which allows for a great control of nutrient availability. This way they can add and remove different nutrients to see how the plant reacts
most plant dry biomass is built from what
the CO2 in air
how are plants able to obtain all their nutrients if they cant move
plant growth is their version of movement. Roots grow through the soil to obtain mineral nutrients and water. Stem and leaf growth helps a plant get more light and CO2
how does the soil direct root growth
different situations can create soil that is more alkaline, or acidic. And such microenvironments encourage or discourage further growth of a root system and help direct its growth
what happens to root growth when phosphorus is deficient
Low phosphorus levels cause the root system in the topsoil to branch out more, as well as grow longer root hairs, where the likelihood of reaching phosphorus rich patches of soil may increase.
what happens to root growth when nitrogen is deficient
Low nitrogen levels cause the root system to grow deeper into the soil where the likelihood of reaching nitrate-rich patches of soil may increase
how are nutrients transported into a cell
ions are actively transported with transport proteins across the cell membrane of epidermal cells into the symplast because their concentrations in the soil solution are generally lower than their concentrations inside the cells.
why is nutrient uptake so highly regulated
the levels of ions inside cells must be maintained at constant levels
how is nutrient uptake regulated
The number of transporters for a given nutrient can be regulated in response to nutrient availability. Low nutrient levels stimulate transcription, whereas high levels repress transcription
functions of soil
mechanical support
mineral nutrients and water from the soil solution
O2 for root respiration
living components of soil
bacteria, fungi, protists, and animals (earthworms and insects)
nonliving components of soil
rock fragments that range in size from large stones to sand to silt, to clay. Soil also contains water and dissolved mineral nutrients, air spaces, and dead organic matter
name the typical percentages of living and nonliving components in soil
particles: 45%
water: 25%
air:25%
organisms:5%
what is the A horizon of soil
A horizon is the topsoil that supports the plant’s nutrient needs. it contains most of soil’s living and dead organic matter

what is the B horizon of soil
B horizon is the subsoil, which accumulates materials from the topsoil above it and from the parent rock below

what is the C horizon of soil
C horizon is the parent rock, also called bedrock, which is in the process of breaking down to form soil

what is soil fertility
a soil’s ability to support plant growth
what factors determine a topsoil’s fertility
their proportions of sand, silt, and clay