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Darwin’s hypothesis
Organisms that left south America to go to Galapagos Islands allowed them to diversify and gave rise to new species.
Descent with modification
Evolution
Change in genetic make up of a population over time; descent with modification
Natural Selection
A process in which individuals who have certain traits tend to survive and reproduce at higher rates than other individuals because of those traits
Phenotypic variations in the population
Favorable phenotypes are measured by fitness through reproductive success
Traits are heritable
More offspring are produced that can survive
Selective pressures
External environmental factors that influence which organisms can survive+reproduce
Adaptations
Inherited characteristics of organisms that enhance their survival and reproduction
Differential survival
Unequal ability of individuals within a population to survive and reproduce due to variations in their inherited traits → competition is the cause
Artificial selection
Selective breeding of domesticated plants and animals to encourage the occurrence of desirable traits
Ethology
The study of how evolutionary processes shape inherited behaviors and the ways that animals respond to specific stimuli
Proximate Cause
How a behavior occurs or how it is modified
What’s the stimulus? More emphasis on nurture than nature
Ultimate Cause
Why a behavior occurs?
nature heavy rather than nurture
Innate behaviors
Developmentally fixed, inherited, born with
Learned behaviors
Depend on environmental influence, experiences affect, high variation in a population
Fixed action patterns (FAPs)
Sequence of unlearned acts directly linked to a stimulus. Innate, stereotypical behavioral sequences, involuntary
actions are unchangeable
carried out to completion
triggered by a sign stimulus (external)
Migration
Innate behavior, a regular long-distance change in location
triggered by environmental cues - sun position, earth’s magnetic field, body clock
Pheromones
Chemicals emitted by members of a species that can affect other membranes of the same species
Stimulus response chains
what a response to a stimulus serves as the next stimulus for a behavior
Waggle Dance
Honey bees communicate through body movements
Directed movements
Movements towards or away from a stimulus
Kinesis
A change in the rate of movement or the frequency of turning movements in response to a stimulus, non-directional
Taxis
Directional movement towards (positive) or away from (negative) a stimulus
Phototaxis
Movement in response to light
Chemotaxis
Movement in response to chemical signals
Geotaxis
Movement in response to gravity
Learning
Modification of behavior based on specific experiences
Imprinting
Long-lasting behavioral response to an individual
during sensitive period of development (early in life)
Occurs on the first individual they encounter (ducks follow their mothers)
A learned behavior!
Spatial learning
establishing memories based upon the spatial structure of the animal’s surroundings
cognitive maps are formed, using landmarks and environmental cues
ex: bird going back to its hidden nest
Associative learning
Ability to associate one environmental feature with another
ex: associating monarch butterflies with a foul taste
Social learning
Learning through observations and limitations of the observed behaviors
ex: chimps breaking open oil palm nuts
Mating bahviors
Animals can be monogamous or polygamous (one partner or multiple)
Altruism
Selfless behavior
reduces fitness of individual but increases fitness of the population
Phototropism
A directional response that allows plants to grow towards (or away from sometimes) a source of light
Photoperiodism
Allows plants to develop in response to day length ; plants flower only at certain times of the year
Gene Pool
A population’s genetic makeup
all copies of every type of allele
if theres only one allele present for a particular locus in the population, its is fixed
Microevolution
Small-scale genetic changes in a population
driven by random occurrence (genetic drift, mutations, gene flow/migration, natural selection)
Genetic drift
Chance events that cause a change in allele frequency from one generation to the next
can lead to loss of genetic variation
can cause harmful alleles to become fixed
two types:
bottleneck effect
founder effect
Bottleneck effect
When a large population is reduced by a non-selective disaster (floods, famine, fire, hurricane, hunting)
Some alleles in the surving population become overrepresented, underrepresented, or absent
Founder effect
When few individuals become isolated from a large population and establish a new small population with a gene pool that differs from the large population
loss of genetic diversity
Gene Flow
The transfer of alleles into or out of a population due to fertile individuals or gametes
ex: pollen being blown to a new location
Directional Selection
Selection towards one extreme phenotype
Stabilizing selection
Selection towards the mean and against the extreme phenotypes
Disruptive selection
Selection against the mean, both phenotypic extremes have the highest fitness
Sexual selection
A type of natural selection that explains why many species have unique/showy traits
Males often have useless structures simply because females chose that trait
Can produce traits that are harmful to survival
ex: colorful features in male peacocks make then easier to spot by predators
***Makes certain inherited traits more successful when trying to mate and reproduce***
Hardy Weinberg Principle
The frequencies of alleles and genotypes in a population will remain constant from generation to generation, provided that only Mendelian segregation and recombination of alleles are at work
Hardy Weinberg Equilibrium
no mutations
random mating
no natural selection
extremely large population size
no gene flow
p + q = 1
p = frequency of the dominant allele in a population
q = frequency of the recessive allele in a population
p²+2pq+q²=1
p²=percentage of the homozygous dominant individuals
pq=percentage of the heterozygous individuals
q²=percentage of the homozygous recessive individuals
Change in population
dN/dt = B - D
B = birth rate
D = death rate
dN/dt = change in population size
Exponential Growth
J shaped curve
dN/dt = r(max)*N
r(max) = max per capita growth rate of the population
N = population size
Logistic Growth
Per capita rate of increase approaches zero as the population size its carrying capacity
dN/dt = r(max)*N*((K-N)/K)
r(max) = maximum per capita growth rate of the population
N = population size
K = carrying capacity
K-selection (density-dependent)
Selection for life history traits that are sensitive to population density
high-density populations that are close to carrying capacity (K)
R-selection (density independent)
Selection for life history traits that maximize reproductive success
seen in low density populations with little competition
Life History
the traits that affect an organism’s schedule of reproduction and survival
when reproduction begins
how often the organism can reproduce
the number of offspring produce per reproductive episode
Density-dependent regulation
factors that can slow or stop growth by decreasing birth rate or increasing death rate in population
competition, predation, toxic wastes, territoriality, disease
Density-independent regulation
factors that exert their influence on population size, but the birth/death rate doesn’t change
weather, climate, natural disasters
Allopatric Speciation
Physical barrier divides population
or Small population is separated from main population
prevents gene flow
Sympatric Speciation
New species evolved while still inhabiting the same geographic region as the ancestral species
→Usually due to exploitation of a new niche
Speciation
Formation of new species due to reproductive isolation:
pre/postzygotic barriers
Prezygotic barriers
Prevent mating or hinder fertilization:
habitat
temporal
behavioral
mechanical
gametic
Habitat isolation
Species live in diff areas or the occupy different habitats within the same area
Temporal Isolation
Species breed at different times of the day, year, or season
Behavioral isolation
Unique behavioral rituals separate species
Mechanical isolation
Reproductive anatomy of one species doesn’t fit that of the other species
Gametic isolation
Proteins on the surface of gametes do not allow for the egg and sperm to fuse
Postzygotic Barriers
Prevent a hybrid zygote from developing into a viable, fertile adult
reduced hybrid viability
reduced hybrid fertility
hybrid breakdown
Reduced Hybrid Viability
The genes of different parent species may interact in ways that impair the hybrid’s (child) development or survival
ex: domestic sheep + domestic goat = hybrid embryo dies early on
Reduced hybrid fertility
A hybrid can develop into a healthy adult, but it is sterile, not reproductive
Usually due to differences in number of chromosomes between parents
Hybrid breakdown
The hybrid of the first gen may be fertile but when they mate with a parent species or one another, their offspring will be sterile
Microevolution
Change in allele frequencies within a single species or population (natural/sexual selection, genetic drift, gene flow)
Macroevolution
Large evolutionary patterns (adaptive radiation, mass extinction)
Punctuated equilibrium
When evolution occurs rapidly after a long period of stasis. Stop-Start-Stop
Gradualism
When evolution occurs slowly over 100s, 1000s, 1000000s of years
Divergent evolution
Groups with the same common ancestor evolve and accumulate differences resulting in the formation of a new species
Adaptive radiation
If a new habitat or niche becomes available, species can diversify rapidly
think of the diff versions of Darwin’s finches
Convergent evolution
Two different species develop similar traits despite having different ancestors due to environmental pressures
analogous traits
Fundamental niche
The niche (role) potentially occupied by the species if there were no limiting factors (predators, competitors, etc)
Realized niche
The portion of the fundamental niche the species actually occupies
Interspecific interactions
Interactions of individuals from one species with individuals of another
competition, predation, herbivory, symbiosis, facilitation
Competitive exclusion principle
Two species competing for the same resource cannot coexist permanently
Niche partitioning
Natural selection drives competing species into different patterns or resource use, or different niches
Cryptic coloration
Camouflage
Batesian mimircy
Harmless species mimics a harmful one
Mullerian mimicry
Two or more bad-tasing species resemble each other
Parasitism
(+/-) when one organism (parasite) derives nourishment from another (host)
Mutualism
(+/+) when both organisms benefit from the relationship
Commensalism
(+/0) when one organism benefits and the other is neither harmed or unharmed, neutral
Facilitation
(+/+) or (+/0) when one species has a positive effect on the survival and reproduction of another without an intimate association of symbiosis
common in plants
Species Richness
number of diff species
Relative abundance
The proportion each species represents of all the individuals in the community
Simpson’s diversity index
Used to calculate diversity based on species richness + rel. abundance
Diversity Index = 1 - sum of(n/N)²
n= total number of organisms of a particular species
N= total number of organisms of all species
High Div. index = high biodiversity